Sous-embranchement (subphylum) des Rhynchonelliformea  
Classe des Rhynchonellata 
Ordre des Terebratulida 

Sous-ordre des Terebratulidina 
Super-famille des Terebratuloidae 
Super-famille des Dyscolioidae 
Super-famille des Cancellothyroidea 

Genus Liothyrella Thomson, 1916
[= Dolichosina Cooper, 1983; Mimorina Cooper, 1983]
[Type species = Terebratula uva Broderip, 1833 (p. 124)]

Large to very large, elongate oval to subcircular; ventribiconvex, anterior commissure usually rectimarginate, occasionally broadly uniplicate; smooth or with faint, radial or zigzag capillae; beak usually short, suberect; foramen usually large, submesothyrid, labiate; symphytium wholly or partly visible; pedicle collar short, teeth triangular, narrow; low myophragm may be present; cardinal process transverse semiellipse; outer hinge plates variable in width and length; attached near dorsal edge of crural bases; loop variable; usually widely triangular, crural processes located near socket openings, transverse band relatively narrow, variable; spicules abundant.

Eocene - Present

Diagnosis from volume 5 of the
Treatise on Invertebrate Paleontology (2006)

Extant Species of Liothyrella
  • L. uva (Broderip, 1833)
  • L. moseleyi (Davidson, 1878)
  • L. winteri (Blochmann, 1906)
  • L. neozelanica Thomson, 1918
  • L. multiporosa Foster, 1974
  • L. delsolari Cooper, 1982
  • Diagnosis


    Liothyrella uva   (Broderip, 1833)

    Type locality: “in sinu Tehuantepec” - “found by Captain Dare, attached to a sea-vorn bivalve, at a depth of 10-12fathoms”.
    Depth range: 0 - 2273 m

    Terebratula uva Broderip, 1833
    Liothyris uva Davidson, 1886
    ?Terebratula uva Gray, 1853
    Terebratula moseleyi Fischer & Œhlert, 1892
    Liothyrella antarctica Blochmann, 1906
    Liothyrina uva Blochmann, 1906
    Liothyrina moseleyi Dall, 1908
    Liothyrina uva var. notorcadensis Jakson, 1912
    Liothyrina antarctica Joubin, 1914
    ?Liothyrella ovata Thomson, 1918
    Liothyrella notorcadensis Thomson, 1918
    Gryphus antarcticus Dall, 1920
    Gryphus uva Dall, 1921
    Liothyrella oblonga Cooper, 1973
    Liothyrella antartica georgina Zezina 1980
    Liothyrella "uva" Cooper, 1983
    Liothyrella uva georgina Foster, 1974
    Liothyrella uva notorcadensis Foster, 1974
    Liothyrella uva cancerderma Foster, 1974
    Liothyris uva Zezina, 1976
    Liothyrella georgina Cooper, 1982
    Liothyrella fosteri Cooper, 1982
    Liothyrella expansa Cooper, 1982
    Liothyrella notocadensis Cooper, 1982


    Diagnose in preparation -



    Liothyrella moseleyi   (Davidson, 1878)

    Type locality: West of Kergelen Island, 210 fathoms [46°47'S, 51° 37’E, depth 384 m]
    Depth range: some meters - 700 m

    Terebratula moseleyi Davidson, 1878
    Liothyris moseleyi Davidson, 1886
    Terebratula (Liothyrina) moseleyi Davidson, 1892
    Liothyrina moseleyi Blochmann, 1906
    Liothyrella moseleyi Hertlein & Grant, 1944

    Diagnosis - in preparation


    Liothyrella winteri   (Blochmann, 1906)

    Type locality: “Valdivia, Station 165, St. Paul Tiefe 680 m [38°40,0'S, 77°38,6'E, depth  672 m, NE of Saint-Paul Island, Valdivia-Expedition, 1898-1899].
    Depth range: 672 - 680 m

    Liothyrina winteri Blochmann, 1906
    Liothyrella winteri Thomson, 1927

    Diagnosis - in preparation


    Liothyrella neozelanica   Thomson, 1918

    Type locality: Cook Strait, off Wellington, 370 m
    Depth range: 6 - 1939 m


    Liothyrina sp. Jackson, 1918
    Liothyris neozelanica Zezina, 1976

    Diagnosis - in preparation


    Should be added:
  • Muscle disposition: see Fig. in front -











    Disposition of the pedicle and pedicles muscles - sagittal section
    (from Richardson, 1979)


    Muscle attachment (from Richardson, 1979)


    L. delsolari  Cooper, 1982

    Type locality: off Peru [4°00'S, 80°30'W]
    Depth range: 760 - 1208 m


    Diagnosis - in preparation