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List of the benthic biocenoses in the Mediterranean Sea

Updated by Christian C. EMIG – April, 28 2008 from the cited references

 

SUPRALITTORAL

 

MUDS

     Biocenosis of beaches with slow-drying wracks under glassworts


SANDS

     Biocenosis of supralittoral sands

       Facies of sands without vegetation, with scattered debris

       Facies of depressions with residual humidity

       Facies of quickly-drying wracks

       Facies of tre truncks which have been washed ashore

       Facies of phanerogams which have been washed ashore (upper part)


STONES and GRAVELS

     Biocenosis of slow-drying wrack


HARD SUBSTRATES and ROCKS

     Biocenosis of supralittoral rock

       Association with Entophysalis deusta and Verrucaria amphibia

       Pools with variable salinity (mediolittoral enclave)

 

MEDIOLITTORAL top

 

MUDS, SANDY MUDS and SANDS

     Biocenosis of muddy sands and muds

       Association with halophytes

       Facies of saltworks


SANDS

     Biocenosis of mediolittoral sands

       Facies with Ophelia bicornis


STONE and GRAVELS

     Biocenosis of mediolittoral coarse detritic bottoms

       Facies of banks of dead leaves of Posidonia oceanica and other phanerogams


HARD SUBSTRATES and ROCKS

     Biocenosis of the upper mediolittoral rock

       Association with Bangia atropurpurea

       Association with Porphyra leucosticta

       Association with Nemalion helminthoiof and Rissoella verruculosa

       Association with Lithophyllum papillosum and Polysiphonia spp

     Biocenosis of the lower mediolittoral rock

       Association with Lithophyllum lichenoides (= Cornice of L. tortuosum)

       Association with Lithophyllum byssoides

       Association with Tenarea undulosa

       Association with Ceramium ciliatum and Corallina elongata

       Facies with Pollicipes cornucopiae

       Association with Enteromorpha compressa

       Association with Fucus virsoides

       Neogoniolithon brassica-florida concretion

       Association with Gelidium spp

       Pools and lagons sometimes associated with vermetids (infralittoral enclave)

     Mediolittoral caves

       Association with Phymatolithon lenormandii and Hildenbrandia rubra

 

INFRALITTORAL top

 

SANDY MUDS, SANDS, GRAVELS and ROCKS in EURYHALINE and EURYTHERMAL ENVIRONMENT

     Euryhaline and eurythermal biocenosis (BEE)

       Association with Ruppia cirrhosa and/or Ruppia maritima

       Facies with Ficopomatus enigmaticus

       Association with Potamogeton pectinatus

       Association with Zostera noltii in euryhaline and eurythermal environment

       Association with Zostera marina in euryhaline and eurythermal environment

       Association with Gracilaria spp

       Association with Chaetomorpha linum and Valonia aegagropila

       Association with Halopithys incurva

       Association with Ulva laetevirens and Enteromorpha linza

       Association with Cystoseira barbata

       Association with Lamprothamnium papulosum

       Association with Cladophora echinus and Rytiphloea tinctoria


FINE SANDS MORE or LESS MUDDY

     Biocenosis of fine sands in shallow waters (SFHN)

       Facies with Lentidium mediterraneum

     Biocenosis of well-sorted fine sands (SFBC)

       Association with Cymodocea nodosa on well sorted fine sands

       Association with Halophila stipulacea

     Biocenosis of superficial muddy sands in sheltered waters (SVMC)

       Facies with Callianassa tyrrhena and Kellia corbuloides

       Facies with freshwater resurgences with Cerastoderma glaucum, Cyathura carinata

       Facies with Loripes lacteus and Tapes spp

       Association with Cymodocea nodosa on superficial muddy sands in sheltered waters

       Association with Zostera noltii on superficial muddy sands in sheltered waters

       Association with Caulerpa prolifera on superficial muddy sands in sheltered waters

       Facies of hydrothermal oozes with Cyclope neritea and nematodes


COARSE SANDS MORE or LESS MUDDY

     Biocenosis of coarse sands and fine gravels mixed by waves

       Association with rhodolithes

     Biocenosis of coarse sands and fine gravels under bottom currents (SGCF) (also found in the Circalittoral)

       Maërl facies (= Association with Lithothamnion corallioides and Phymatolithon calcareum) (can also be found as facies of the biocenosis of coastal detritic)

       Association with rhodolithes


STONES and GRAVELS

     Biocenosis of infralittoral pebbles

       Facies with Gouania wildenowi

     Biocenosis of Posidonia oceanica meadow (HP) (= Association with Posidonia oceanica)

       Ecomorphosis of stripped meadows

       Ecomorphosis of « barrier reef » meadows

       Facies of dead « mattes » of Posidonia oceanica

       Association with Caulerpa prolifera


HARD SUBSTRATES and ROCKS

     Biocenosis of infralittoral algae (AP)

       Overgrazed facies with encrusting algae and sea urchins

       Association with Cystoseira amentacea (var. amentacea, var. stricta, var. spicata)

       Facies with vermetids

       Facies with Mytilus galloprovincialis

       Association with Corallina elongata and Herposiphonia secunda

       Association with Corallina officinalis

       Association with Codium vermilara and Rhodymenia ardissonei

       Association with Dasycladus vermicularis

       Association with Alsidium helminthochorton

       Association with Cystoseira tamariscifolia and Saccorhiza polyschides

       Association with Gelidium spinosum hystrix

       Association with Lobophora variegata

       Association with Ceramium rubrum

       Facies with Cladocora caespitosa

       Association with Cystoseira brachycarpa

       Association with Cystoseira crinita

       Association with Cystoseira crinitophylla

       Association with Cystoseira sauvageauana

       Association with Cystoseira spinosa

       Association with Sargassum vulgare

       Association with Dictyopteris polypodioides

       Association with Calpomenia sinuosa

       Association with Stypocaulon scoparium (=Halopteris scoparia)

       Association with Trichosolen myura and Liagora farinosa

       Association with Cystoseira compressa

       Association with Pterocladiella capillacea and Ulva laetevirens

       Facies with large hydrozoans

       Association with Pterothamnion crispum and Compsothamnion thuyoides

       Association with Schottera nicaeensis

       Association with Rhodymenia ardissonei and Rhodophyllis divaricata

       Facies with Astroides calycularis

       Association with Flabellia petiolata and Peyssonnelia squamaria

       Association with Halymenia floresia and Halarachnion ligulatum

       Association with Peyssonnelia rubra and Peyssonnelia spp

       Facies and association of the coralligenous biocenosis (in enclave)

 

CIRCALITTORAL top

 

MUDS

     Biocenosis of coastal terrigenous muds (VTC)

       Facies of soft muds with Turritella tricarinata communis

       Facies of sticky muds with Virgularia mirabilis and Pennatula phosphorea

       Facies of sticky muds with Alcyonium palmatum and Stichopus regalis


SANDS

     Biocenosis of the muddy detritic bottom (DE)

       Facies with Ophiothrix quinquemaculata

     Biocenosis the coastal detritic bottom (DC)

       Association with rhodolithes

       Maerl facies (Lithothamnion corallioides and Phymatholithon calcareum)

       Association with Peyssonnelia rosa-marina

       Association with Arthrocladia villosa

       Association with Osmundaria volubilis

       Association with Kallymenia patens

       Association with Laminaria rodriguezii on detritic

       Facies with Ophiura texturata

       Facies with Synascidies

       Facies with large bryozoans

     Biocenosis of coarse sands and fine gravels under the influence of bottom currents (SGCF) (can be found in the Infralittoral)


HARD SUBSTRATES and ROCKS

     Coralligenous biocenosis (C)

       Association with Cystoseira zosteroides

       Association with Cystoseira usneoides

       Association with Cystoseira dubia

       Association with Cystoseira corniculata

       Association with Sargassum spp

       Association with Mesophyllum lichenoides

       Association with Lithophyllum frondosum and Halimeda tuna

       Association with Laminaria ochroleuca

       Association with Rodriguezella strafforelli

       Facies with Eunicella cavolinii

       Facies with Eunicella singularis

       Facies with Lophogorgia sarmentosa

       Facies with Paramuricea clavata

       Facies with Parazoanthus axinellae

       Facies with Parazoanthus axinellae

     Semi-dark caves (also in enclave in the Infralittoral)

       Facies with Corallium rubrum

       Facies with Leptosammia pruvoti

 

BATHYAL top

 

MUDS

     Biocenosis of muds of bathyal superior (VBS) (hypothetic)

     Biocenosis of (bathyal) deep-sea muds (VP)

       Facies of sandy muds with Thenea muricata

       Facies of fluid muds with Brissopsis lyrifera

       Facies soft muds with Funiculina quadrangularis and Apporhais seressianus

       Facies of compact muds with Isidella elongata

       Facies with Pheronema grayi


SANDS

     Biocenosis of shelf-edge detritic bottom (DL)

     Biocenosis of bathyal detritic sands (SDB)

       Facies of Neolampas rostellata


HARD SUBSTRATES and ROCKS

     Biocenosis of offshore rocks (RL)

     Biocenosis of deep-sea (white) corals (CB)

       Facies of Errina aspera and Pachylasma giganteum

     Community of deep-sea cold-vent (hydrothermal vent)

 

ABYSSALtop

 

MUDS

     Biocenosis of the abyssal mud (VA)