◄ Carnets Geol. 15 (13) ►
Contents
[Introduction]
[Pseudohaploceras liptoviense (, 1856)]
[Deshayesites sp.]
[Dufrenoyia sp.] [Age assignment]
[Conclusions] and ... [Bibliographic references]
Instituto de Geología, Universidad Nacional Autónoma de México, Ciudad
Universitaria, Coyoacán, 04510, México, Distrito Federal (Mexico)
Tel.: +52 55 5622 4280 ext. 156
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76118, Texas (United States of America)
Published online in final form (pdf) on August 6, 2015
[Scientific
editor:
Michel ; technical editor: Bruno ;
language editor: Stephen ]
We study the three best preserved ammonites collected by in 1924-1925 in Lleida Province, Catalonia, Spain, in order to assess their taxonomic assignment. We also include images of these ammonoids since they were not illustrated in 's original 1934 work. This taxonomic review allows us to identify these lower Aptian ammonites as Pseudohaploceras liptoviense, Deshayesites sp. and Dufrenoyia sp.
Ammonites; ; Aptian; Catalonia; Spain.
J.A. & K.P. (2015).- Review of some Aptian ammonites collected by Gaston in Lleida Province, Catalonia, Spain.- Carnets Geol., Madrid, vol. 15, nº 13, p. 173-178.
Révision de quelques ammonites aptiennes récoltées par Gaston dans la Province de Lérida (Catalogne, Espagne).- Nous avons étudié les trois ammonites les mieux préservées parmi celles récoltées par en 1924-1925 dans la Province de Lérida (Catalogne, Espagne) afin d'en ré-évaluer plus sûrement l'attribution taxinomique. Nous figurons ces ammonoïdés puisqu'il n'y avait aucune illustration dans la publication originale de en 1934. Ce ré-examen taxinomique nous permet d'attribuer ces trois ammonites de l'Aptien inférieur à Pseudohaploceras liptoviense, Deshayesites sp. et Dufrenoyia sp.
Ammonites ; ; Aptien ; Catalogne ; Espagne.
Revisión de algunos ammonites aptienses colectados por Gaston en la Provincia de Lérida, Cataluña, España.- Se estudian los tres ammonites mejor conservados colectados por en 1924-1925 en la Provincia de Lérida, Cataluña, España, con la finalidad de mejorar su asignación taxonómica. También incluimos imágenes de estos ammonites ya que no se figuraron en el trabajo original de , 1934. Esta revisión taxonómica nos permite identificar los ammonites del Aptiense inferior: Pseudohaploceras liptoviense, Deshayesites sp. y Dufrenoyia sp.
Ammonites; ; Aptiense; Cataluña; España.
Revisió d'alguns ammonits aptians recollits per Gaston a la Província de Lleida, Catalunya, Espanya.- S'estudien els tres ammonits millor conservats recollits per en 1924-1925 a la Província de Lleida, Catalunya, Espanya, per tal d'avaluar la seva assignació taxonòmica. També s'inclouen imatges d'aquests ammonits ja que no s'il·lustren en l'obra original d' del 1934. Aquesta revisió taxonòmica ens permet identificar aquests ammonits de l'Aptià inferior com: Pseudohaploceras liptoviense, Deshayesites sp. i Dufrenoyia sp.
Ammonits; ; Aptià; Catalunya; Espanya.
In 1922, under the supervision of Charles and Paul , Gaston began his doctoral studies of the geology along the south side of the Pyrenees (, 2010, p. 66; & , 2010, p. 28). During his field work in 1924 and 1925, he found some Aptian ammonites in the Segre River region, Lleida Province, Catalonia. Although he never completed his PhD (, 2010, p. 66), in 1934 he published a short paper describing two of these ammonites. However, he did not include any illustrations or figures to accompany his detailed descriptions and comparisons with closely related taxa. Fortunately, these specimens are still housed in the collections of the Université Paul Sabatier of Toulouse, France (= UPS collection). We were able to locate 's published material, as well as two additional unpublished specimens. In this work, we review these four ammonites, three of which are illustrated for the first time. Our reexamination of these ammonites sheds light on the work of (1934) and contributes to the general knowledge of the Aptian ammonites of Catalonia. Recently in one area of Lleida Province, the Organyà Basin, the stratigraphy, sedimentology and geochemical record of the uppermost Barremian-lowermost Aptian hemipelagic sediments has been investigated ( & , 2014; et al., 2014). The current study of the ammonoid record will be useful in establishing a precise age determination of these hemipelagic sediments. It is part of a broader examination of the ammonoid record of the uppermost Barremian-Aptian of Lleida Province.
(1934, p. 223) identified the first specimen as Puzosia
liptoviensis (Fig. 1.A 
,
UPS-GA1934-01-BP-7), collected in 1924 by himself
between Gosol and la Collada de Molas. This taxon is currently assigned to the
genus Pseudohaploceras as Pseudohaploceras
liptoviense (, 1856). P.
liptoviense has a distinctive rib pattern that consists of small, flexuous,
fold-like primary and secondary ribs, interrupted at irregular intervals by
bifurcations and much larger primaries that are frequently paired with
constrictions (see et al., 1996, Fig. 52, 3a). In P.
liptoviense, the larger primaries are often strongly pronounced towards the
umbilicus, especially on the juvenile whorls ( & ,
2011, Pl. 1, figs. 1-5). Due to the poor preservation of the specimen, its
identification is difficult. However, 's original
assignment seems to
be correct. The irregularity in the rib pattern separates P. liptoviense from other similar species, such as P.
matheroni (, 1841), which has a more regular rib pattern,
consisting of groups of 8-14 finer ribs, interrupted by larger, cord-like
primaries. In P. matheroni, these larger ribs are frequently paired with one wide
and one narrow constriction on each side. The
robust costation of the specimen studied here also allows us to distinguish 's
specimen from the more weakly ornamented P.
douvillei (, 1920) (see: ,
1992, Pl. 4, figs. 1-6;
et al., 2006, p.
53-54).
P. ramosum (, 1991) is
a more closely related taxon that belongs to the P. liptoviense group ( & ,
2004). The differences between the two taxa are subtle, P. ramosum having fewer
secondary ribs and thicker whorls. Another species that shows some similarities
is "Pseudohaploceras" angladei
(, 1890) but this ammonite has a thicker whorl section, which may
even be depressed, and probably belongs to the genus Valdedorsella.
Click on thumbnail to enlarge the image.
Figure 1: A. Pseudohaploceras liptoviense (, 1856) (= Puzosia liptoviensis .- , 1934), lateral view of UPS-GA1934-01-BP-7. B. Deshayesites sp. (= Ammonitoceras cf. Lahuseni .- , 1934), lateral view of UPS-GA1934-02. C. Dufrenoyia sp. (Parahoplites furcatus (Type Dufrenoyi).- 's specimen label), lateral view of UPS-GA1934-03. Scale bar is 10 mm.
's
specimen (Fig. 1.B , UPS-GA1934-02) is a fragment of a large shell.
(1934, p. 223) wrote that this fragment is 17 cm long and 9 cm high and
suggested that the ammonite could have been as large as 30-40 cm in diameter.
The fragmentary ammonite has a tall, elliptical whorl section with a rounded
ventral region (Fig. 2.A ). The costation consists of slightly sigmoidal primary
ribs, accompanied by scarce secondary ribs that can be intercalated or
bifurcated about the middle of the flank.
Ammonitoceras lahuseni
has a circular to depressed whorl section as does any other species of Ammonitoceras
( et al., 1996, p. 221; & ,
2011,
Figs. 1-2).
This important feature allowed us to rule out the assignment of the fragmentary
specimen to the genus Ammonitoceras.
commented on this apparent difference, comparing the whorl section of his
specimen to those of Ammonitoceras ucetiae
, 1876, which were collected in Santander, Cantabria, northern
Spain by (1920, p. 140) and identified by Wilfrid .
(1934) commented that the whorl section of this specimen is very similar to that
of 's material. However, we also have access to two
gypsum casts of 's specimens. One of them
is slightly crushed, having a whorl section similar to 's
specimen, but the second uncrushed specimen clearly has a circular whorl
section. In our opinion, 's specimen is not crushed, and its whorl
section matches that of Deshayesites
but certainly not that of Ammonitoceras (Fig. 2 ). The rib pattern of the ammonoid studied here is consistent with the
ontogenic development of Deshayesites,
where the progressive loss of secondary ribs and slight straightening of both
the primaries and the secondaries are particularly characteristic of the adult
macroconch specimens (, 1964, Pl. 43, fig. 1a, and Pl. 54, fig. 1a;
et al., 2010, electronic appendix Fig. VC). In addition the large size of the
fragment suggests that it belongs to an adult macroconch of Deshayesites.
Based on the weight of evidence, we conclude that this ammonite is a fragment of
an adult macroconch of Deshayesites,
which we report here as Deshayesites
sp., since there is not enough information to make a specific determination.
Click on thumbnail to enlarge the image.
Figure 2: A. Whorl section of Deshayesites sp. (= Ammonitoceras cf. Lahuseni .- , 1934), specimen UPS-GA1934-02. B. Whorl section of the holotype of Ammonitoceras ucetiae , 1876, type species of the genus Ammonitoceras. Modified from Figure 2 of and (2011). Scale bar is 10 mm.
At
the end of his paper, (1934, p. 225) wrote that he also found
better known and arguably less important species of cephalopods in Lleida
Province, including ammonites similar to Parahoplites
furcatus and Parahoplites dufrenoyi (=
Dufrenoyia spp.). We have found
another partially crushed specimen at Organyà, Lleida Province (Fig.
1.C ,
UPS-GA1934-03), the third one examined here, labeled as "Parahoplites furcatus (Type Dufrenoyi)" (Fig. 3 ).
The ornamentation of this specimen consists of paired sigmoidal primary and
secondary ribs (Fig. 1.C ), which are typical of the rib pattern of Dufrenoyia.
However, an abrupt thinning of the whorl is also present
along the last third of the flank and ventral margin, which is typical of
crushed specimens of Dufrenoyia. We
therefore conclude that this is a specimen of Dufrenoyia.
This genus is characterized by a costal section with a flat venter and by the
interruption of the ribs across the midline in early ontogenetic stages. The
more common European species Dufrenoyia
furcata (, 1836) and Dufrenoyia
dufrenoyi (, 1840), can be distinguished because in D.
furcata the ribs cross the venter earlier during ontogenic development (
& , 2010). Since the venter of UPS-GA1934-03 is not
preserved well enough to make a specific determination, we report it here as Dufrenoyia sp.
In the UPS Collection we also found a very small pyritic specimen (6 mm max. diam.) without ribs that was labeled as "Parahoplites furcatus". In our opinion, it is impossible to identify this ammonite, even at the generic level.
Click on thumbnail to enlarge the image.
Figure 3: Original label of Dufrenoyia sp., UPS-GA1934-03.
Deshayesites and Dufrenoyia are index genera of the lower Aptian. Deshayesites ranges through almost all of the lower Aptian, except the uppermost part, at which point the genus Dufrenoyia first appears and replaces Deshayesites. The specimen of Deshayesites sp. collected by is too poorly preserved to place it securely within the biostratigraphical divisions founded on genus Deshayesites within the lower Aptian.
Dufrenoyia, in turn, characterizes the Dufrenoyia furcata Zone and disappears at the lower-upper Aptian transition. (1934, p. 221) reported his Pseudohaploceras liptoviense from the Gargasian, an old term that has an ambiguous chronostratigraphic meaning today, but which encompasses strata of the uppermost lower Aptian to the lowermost upper Aptian in southeast France. This age corresponds to the Dufrenoyia furcata (lower Aptian) and Epicheloniceras martini (upper Aptian) ammonite zones of the standard ammonite zonation of et al. (2011, 2014). However, Pseudohaploceras disappears at the lower-upper Aptian transition, and is succeeded by the upper Aptian desmoceratid Caseyella ( et al., 2012). We conclude then that all of 's specimens reviewed here are lower Aptian without further refinement.
Our review of the three best preserved specimens collected by in 1924-1925 allows us to identify and illustrate for the first time the ammonoid taxa, Pseudohaploceras liptoviense, Deshayesites sp. and Dufrenoyia sp., all of which are early Aptian in age.
This contribution will facilitate continuing study of the Aptian ammonoid record of the Lleida Province.
We appreciated the assistance provided by Michel and access to facilities granted by the UPS to review the ammonites housed in the Université Paul of Toulouse, as well as their help in obtaining certain references. We are very grateful for the helpful corrections and suggestions made by Miguel , Gérard , Antoine and Yves . We are also very grateful to the Language Editor, Stephen , for his corrections that permitted significant improvements to the manuscript.
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