Carnets Geol. 19 (13)  

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Outline:

[1. Introduction] [2. Material and methods]
[3. Systematic palaeontology: Part 1; Part 2; Part 3]
and ... [Bibliographic references]


The bryozoan collection of Prof. Dr Ehrhard Voigt (1905–2004)
at the Senckenberg Institute in Frankfurt.

Part 1 - Introduction and Cyclostomata

Silviu O. Martha

Senckenberg Forschungsinstitute und Naturmuseen, Sektion Marine Evertebraten III (Bryozoologie), Senckenberganlage 25, 60325 Frankfurt am Main (Germany)

Kei Matsuyama

Former address: Senckenberg am Meer, Abteilung Meeresforschung, Südstrand 40, 26382 Wilhelmshaven (Germany)

Joachim Scholz

Senckenberg Forschungsinstitute und Naturmuseen, Sektion Marine Evertebraten III (Bryozoologie), Senckenberganlage 25, 60325 Frankfurt am Main (Germany)

Paul D. Taylor

Departments of Earth and Life Sciences, Natural History Museum, Cromwell Road, London SW7 5BD (UK)

Gero Hillmer

Institut für Geologie, Universität Hamburg, Bundesstr. 55, 20146 Hamburg (Germany)

Published online in final form (pdf) on October 10, 2019
DOI 10.4267/2042/70493

[Editor: Bruno Granier]

Click here to download the PDF version!

Abstract

The bryozoan collection of Prof. Dr Ehrhard Voigt (1905–2004) at the Senckenberg Research Institute in Frankfurt am Main, Germany is a world-renowned collection of great scientific value. It is the world's largest collection of fossil bryozoans from the Upper Cretaceous and Paleocene and a unique archive documenting the evolution of this phylum of marine invertebrates during this time interval in the Boreal Chalk Sea that extended from the British Isles to the Aral Sea in Central Asia. 

The Voigt Collection contains over 300,000 specimens and was relocated to the Senckenberg Institute in 2005 according to the bequest of Ehrhard Voigt. As a result of a DFG-funded project, we present here a three-part type catalogue of the holotypes and neotypes of 256 bryozoan species in the Voigt Collection, of which this is Part 1. In total over the three parts, 247 species are re-illustrated but the name-bearing type specimens of 20 species are missing and no material could be found for 9 species. Two species, described as ctenostome bryozoans by Ehrhard Voigt, are questionable, while a further three 'ctenostome' species and one 'ctenostome' genus are considered as ichnotaxa.

Key-words

• Bryozoa;
• Cheilostomata;
• Cyclostomata;
• Ctenostomata;
• ichnofossils;
• type catalogue;
• palaeontological collections;
• Cretaceous

Citation

Martha S.O., Matsuyama K., Scholz J., Taylor P.M. & Hillmer G. (2019).- The bryozoan collection of Prof. Dr Ehrhard Voigt (1905–2004) at the Senckenberg Institute in Frankfurt. Part 1 - Introduction and Cyclostomata.- Carnets Geol., Madrid, vol. 19, no. 13, p. 221-275.

Résumé

La collection de bryozoaires du Prof. Dr Ehrhard Voigt (1905–2004) conservée à l'Institut Senckenberg de Francfort - 1ère partie - Introduction et Cyclostomes.- La collection de bryozoaires du Prof. Dr Ehrhard Voigt (1905–2004) conservée à l'Institut de recherche Senckenberg de Francfort-sur-le-Main (Allemagne) est une collection de renommée mondiale et de grande valeur scientifique. C'est la plus grande collection au monde de bryozoaires fossiles du Crétacé supérieur et du Paléocène. Elle constitue un patrimoine unique documentant l'évolution de ce phylum d'invertébrés marins au cours de cette période dans la mer de la Craie Boréale qui s'étendait des îles britanniques à la mer d'Aral en Asie centrale. 

La collection Voigt qui compte plus de 300.000 spécimens a été transférée en 2005 à l'Institut Senckenberg grâce à un leg testamentaire d'Ehrhard Voigt. À l'issue d'un programme financé par la DFG, nous présentons ici en trois parties un catalogue des holotypes ou néotypes de quelques 256 espèces de bryozoaires de la collection Voigt, catalogue dont la présente contribution est la première des trois. 247 espèces sont illustrées ici à nouveau, tandis qu'aucun matériel n'a pu être retrouvé pour neuf d'entre elles. Les spécimens-types porte-noms de vingt espèces n'ont pas été retrouvés, Deux espèces, décrites comme des bryozoaires cténostomes par Ehrhard Voigt, sont douteuses, tandis que trois autres espèces "cténostomes" et un genre "cténostome" sont considérés comme des ichnotaxa.

Mots-clefs

• Bryozoa ;
• Cheilostomata ;
• Cyclostomata ;
• Ctenostomata ;
• ichnofossiles ;
• catalogue des types ;
• collections de paléontologie ;
• Crétacé


1. Introduction

Prof. Dr Ehrhard Voigt (1905–2004) was one of the most famous German palaeontologists of the 20th century. His main expertise concerned marine invertebrates of the phylum Bryozoa Ehrenberg, 1831. He began studying Bryozoa as a teenager and went on to publish on bryozoans for almost 80 years (Voigt, 1923; Ernst & Voigt, 2002). Several additional bryozoan projects were completed and published after his death by co-authors, and some unfinished projects are still pending publication. During his lengthy scientific career, Voigt described more than 500 new bryozoan species and subspecies and established 80 genera or subgenera (excluding Foraripora Voigt & Soule, 1973, see 'Remarks for Foraripora pesavis Voigt & Soule, 1973') and 4 families within the phylum Bryozoa. This makes him one of the most prolific and significant bryozoologists.

Voigt's passion and devotion for fossil Bryozoa is legendary among palaeontologists. Encouraged by his parents, he started collecting fossil bryozoans as a child after finding some bryozoans in Upper Cretaceous flints in his parents' garden. As a teenager, he began corresponding with some of the leading bryozoologists of the time and before leaving school, he had already submitted some substantial articles to well-known scientific journals. Given his prowess as a collector, it is not a surprise that Ehrhard Voigt accumulated one of the world's largest collections of fossil bryozoans. Following the loss of his entire collection in 1943 during World War II (Martha, 2014), Voigt amassed an even larger collection. This collection consists of over 300,000 specimens of mostly Upper Cretaceous to Paleocene bryozoans. The material was collected from hundreds of bryozoan-rich exposures in Europe, northern Africa, Central Asia and North America. The focus of the Voigt Collection, however, comprises bryozoans from the Boreal Chalk Sea that endured for almost 50 million years, from the early Cenomanian to the Paleocene, and extended from the British Isles in Western Europe to the Aral Sea in Central Asia. The Voigt Collection represents a unique documentation of the evolution of bryozoans during this time span and is a key resource for anyone seeking to understand the evolutionary history of bryozoans.

The Voigt Collection originally belonged to the Geologisches-Paläontologisches Institut (GPI) at the University of Hamburg, but was relocated to the Senckenberg Forschungsinstiute und Naturmuseen (SGN) in Frankfurt am Main in February 2005 according to the will of Ehrhard Voigt. A DFG project (SCHO 581/12-1 "Enhancing documentation and digitalization of the Bryozoa collection donated by Professor Voigt (1905–2004) to the Senckenberg Research Institute, Frankfurt am Main, Germany") resulted in the digitization of almost 20,000 specimens from the Voigt collection. The top priority of the project was the digitization of the published originals. Thus, 2,666 specimens examined in the works of Ehrhard Voigt and other researchers were found and digitized. Unfortunately, c. 15 % of the originals could not be traced and might be among the thousands of specimens that have not yet been digitized. The digitized data are accessible online at https://search.senckenberg.de/aquila-public-search/search.

Resulting from the DFG project, we present here an illustrated catalogue of the type material from the Voigt Collection lodged at the SGN. During the course of this work, it was discovered that some of the bryozoan material from the Voigt Collection had remained at the GPI in Hamburg where it is now part of the Geologisch-/Paläontologisches Museum (GPM) of the GPI. This material includes all original specimens in the works of Toots (1952), Wiesemann (1963), and Schubert (1986), Voigt's former students, that had previously been part of the Voigt Collection. Furthermore, several specimens described by Voigt (1957a, 1959a), including seven neotypes designated in these works, and from other publications are located in the GPM. Unfortunately, the holotype of Arachnidium jurassicum Voigt, 1977, that should also be in the possession of the GPM could not be found.

The Voigt Collection at the SGN should contain 256 holotypes and neotypes, 66 of which are also name-bearing types of the type species of nominal genera. The holotypes of 20 species (7.8 % of all holotypes and neotypes) could not be found and have to be considered lost (Fig. 1 ). Two species, Amathia immurata Voigt, 1972, and Harmeriella cretacea Voigt, 1957, are of questionable validity. Another three species – Foraripora pesavis Voigt & Soule, 1973, Iramena gosaviensis (Voigt & Soule, 1973), and Pinaceocladichnus echinicola (Voigt & Soule, 1973) – are considered to be ichnospecies, and the genus Foraripora Voigt & Soule, 1973, is an ichnogenus.

Initially, we had intended to include in this type catalogue information and images for all Voigt holotypes and neotypes in the collections of other institutions. We received images from Voigt types lodged at the following institutions:

Unfortunately, as we could not obtain information and images of Voigt types from all collections, our survey remained incomplete. As a consequence, this type catalogue focuses only on the holotypes and neotypes from the Voigt Collection at the Senckenberg Institute in Frankfurt am Main, Germany.

It was found during the work on this type catalogue that many of the species described by Voigt in his earlier publications required revision, beyond the scope of this type catalogue, and we have avoided erecting new genera. Furthermore, since no proper revision of the species is provided, we have avoided the designation of neotypes for the 20 species for which the holotypes could not be found. This is in accordance with Article 75 of the International Code on Zoological Nomenclature (ICZN, 1999).

Fig. 01
Click on thumbnail to enlarge the image.

Figure 1: Bar graph showing the number of missing and available holotypes and neotypes from the Voigt Collection lodged at the Senckenberg Institute, Frankfurt.

On the citation of volume 5 of the Paléontologie française

The Paléontologie française by Alcide d'Orbigny (1802–1857) was published in several releases of a few text pages that were often accompanied by a number of plates (e.g., Fischer, 2002). The publication dates of the text and the plates had long been ambiguous until Sherborn (1899) re-constructed the publication dates of the text livraisons of the Paléontologie française using information from Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde on new literature published. Unfortunately, Sherborn (1899) did not indicate publication dates of the plates accompanying the text livraisons. The lists of new literature published included in Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde, however, included full information on the plates published along with the text livraisons.

Publication dates for the plates of volume 5 of the Paléontologie française containing the bryozoans appear to first have been indicated in Gregory (1909) using the information from Neues Jahrbuch für Mineralogie, Geognosie, Geologie und Petrefaktenkunde. The publication dates have later been used and indicated in reference lists by many other bryozoologists throughout the 20th century (e.g., Walter, 1970; Hillmer, 1971; Brood, 1972). Ehrhard Voigt appears to have not been aware of the publication dates of the Paléontologie française in his very first publications (e.g., Voigt, 1924a, 1924b), but he evidently knew both the publication dates of the text livraisons and the plates as early as 1930, which is evident by comparing the publication dates for Orbigny taxa indicated in Voigt (1930) with the actual publication dates of the text livraisons and the plates. However, he usually cited the fifth volume of the Paléontologie française simply as 'd'Orbigny (1851-1854)' but indicated the publication dates for the plates and text only once in Voigt and Eiserhardt (1995).

The publication dates of at least the plates seem to recently have passed into oblivion, which led to mis-citations of Orbigny's taxa. However, knowledge of the publication dates of the text and the plates of the Paléontologie française has a big effect on dates of publication of the taxa proposed by Orbigny and the first valid name used for a taxon. Article 12.2.7 of the International Code on Zoological Nomenclature considers genus- and species-group names in association with an illustration valid for works published before 1931 (ICZN, 1999). For example, the type species of the cheilostome genus Escharifora Orbigny, 1852, is Escharella argus Orbigny, 1851, and not Escharifora argus Orbigny, 1852, as Pl. 666 of the atlas had already been published one year before publication of p. 209 of the text volume.

The known publication dates of the fifth volume of the Paléontologie française used herein and previously reported by Gregory (1909), Walter (1970), and others are as follows:

2. Material and methods

All material studied and imaged herein is type material (holotypes and neotypes) belonging to the bryozoan collection of Ehrhard Voigt, now lodged at the Senckenberg Forschungsinstitute und Naturmuseen (SGN) in Frankfurt am Main, Germany. All specimens were collected by Ehrhard Voigt in the 1940s to 1990s unless indicated otherwise in the remarks for each species. The neotypes chosen by Voigt (1959a) from material collected by Friedrich von Hagenow (1797–1865) in the middle of the 19th century are now kept together with the Voigt Collection at the SGN.

Macrophotographs were taken using a Keyence 2000D optical microscope at the Senckenberg am Meer, Wilhelmshaven, Germany (SMW). Backscattered electron images of uncoated samples were produced using a Tescan Vega 3 scanning electron microscope equipped with a low-vacuum chamber at the SMW. Material that had already been coated was imaged at the SGN using a Camscan CS 24 scanning electron microscope producing secondary electron images. This material was re-coated (gold/palladium 20:80) before analysis.

For the synonymy lists, the recommendations of Matthews (1972) are partly used. The following signs and abbreviations are used here:

* in front of the year: First valid mentioning of a species name under Article 11 of the International Code on Zoological Nomenclature (ICZN, 1999);

non in front of the year: all material published in this publication is excluded from the species;

p in front of the year: partim, part of the material published in this publication is excluded from the species;

v in front of the year: vidimus, the material used in this publication has been checked by the authors;

? in front of the year: the allocation of this material to the species is questionable because of the way it was presented in that publication.

3. Systematic palaeontology

Phylum Bryozoa Ehrenberg, 1831

Class Stenolaemata Borg, 1926

Order Cyclostomata Busk, 1852

Suborder Articulina Busk, 1859

Family Crisiidae Johnston, 1838

Genus Crisidia Milne Edwards, 1838

Crisidia lagaaiji Voigt, 1987

(Fig. 2a–b )

*# 1987a Crisidia lagaaiji n.sp. – Voigt, p. 13, Pl. 1, figs. 1–10.

Holotype: SMF 25533 (Voigt, 1987a, Pl. 1, fig. 1).

Original label: Voigt collection number 7728 I.

Locus typicus: Caillaud quarry near Ciply, Mons municipality, Wallonia, Belgium.

Stratum typicum: Pockets in the hardground in the basal beds (Tuffeau de la Malogne) of the Tuffeau de Ciply.

Stratigraphical range: Danian.

Further distribution: Danian, Mons borehole, Mons, Wallonia, Belgium; Albert Canal near Riemst-Vroenhoven, Flanders, Belgium; Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands; Sophia Jacoba shaft near Hückelhoven, North Rhine-Westphalia, Germany.

Remarks: The holotype of Crisidia lagaaiji is a single autozooid that is placed together with over 30 other autozooids on a gold foil, coated, and covered by a protective foil. We did not remove the protective foil in order not to damage or lose the holotype. The figures of the colony segments are flipped horizontally in Voigt (1987a).

Suborder Cancellata Gregory, 1896

Family Crisinidae Orbigny, 1853

Genus Polyascosoecia Canu, 1920

Polyascosoecia pseudolichenoides Voigt, 1987

(Fig. 2c–d )

# 1941 Crisina sp. – Hiltermann, Pl. 8, fig. 13.

p# 1972 Crisina lichenoides (Goldfuss), 1827Brood, p. 379, Pl. LIV, figs. 2–4, 7, 11, Pl. LXI, figs. 9, 13.

# 1984 Polyascosoecia pseudolichenoides n.sp. (nom. nud.) – Voigt, Pl. 5, figs. 6–9.

*# 1987a Reteporidea pseudolichenoides n.sp. – Voigt, p. 32, Pl. 20, figs. 5–15.

# 2005 Polyascosoecia pseudolichenoides Voigt, 1984Hinz-Schallreuter & Schallreuter, p. 548.

Holotype: Not found (Voigt, 1987a, Pl. 3, figs. 6–8).

Original label: Voigt collection number 7101 or 7751.

Locus typicus: Glacial drift deposit near Neu Wulmstorf-Daerstorf, Lower Saxony, Germany.

Stratum typicum: Glacial drift deposits.

Further distribution: Danian, Mons borehole and Ciply, Mons municipality, Wallonia, Belgium; Albert Canal near Riemst-Vroenhoven, Waterschei borehole in Genk and Eisden shaft near Maasmechelen, Flanders, Belgium; Curfs Quarry near Berg, Valkenburg aan de Geul municipality; Beatrix and Neer boreholes near Neer, Leudal municipality (all Limburg, Netherlands); Baltic Danian in Denmark and southern Sweden and several glacial drift deposits of Danian age in northern Germany.

Stratigraphical range: Danian.

Remarks: In the species description (Voigt, 1987a), 7101 is indicated as the number of the holotype, while the figure caption gives 7751 as the holotype. The species had been previously mentioned and figured as "Polyascosoecia pseudolichenoides n.sp. (nom. nud.)" by Voigt (1984, p. 398). Although Hinz-Schallreuter and Schallreuter (2005) ascribed the authorship of the species to Voigt (1984) as it fulfilled the provisions of the International Code on Zoological Nomenclature, it is evident that Voigt (1984) did not intend to erect a new species and explicitely considered Polyascosoecia pseudolichenoides as a nomen dubium at that time. The species was formally erected by Voigt (1987a) as Reteporidea pseudolichenoides. However, it was transferred to the genus Polyascosoecia Canu, 1920, by Voigt in a footnote during the printing phase. The holotype was listed by Eiserhardt (1998) as Voigt collection number 7751.

Genus Reteporidea Orbigny, 1849

Reteporidea georgiknetschi Voigt, 1984

(Fig. 2e–f )

?p# 1881 Retepora lichenoides Goldf.Quenstedt, p. 347, Pl. 155, fig. 31 (non Pl. 155, fig. 30).

*# 1984 Reteporidea georgiknetschi n.sp. – Voigt, p. 398, Pl. 3, figs. 1–6, Pl. 4, figs. 1–10, Pl. 5, figs. 1–3.

Holotype: SMF 25297 (Voigt, 1984, Pl. 3, fig. 4).

Original label: Voigt collection number 9796.

Locus typicus: Blom Quarry near Terblijt, Valkenburg aan de Geul municipality, Limburg, Netherlands.

Stratum typicum: Belemnitella junior belemnite Zone, Tuffeau de Maastricht.

Stratigraphical range: Late Maastrichtian.

Further distribution: Late Maastrichtian, Tuffeau de Maastricht, Limburg, Netherlands.

Remarks: In the species description, Voigt (1984, p. 398) mistakenly indicated that the holotype was depicted in Pl. 4, fig. 4. However, the holotype is depicted in Pl. 3, fig.4, while Pl. 4, fig. 4 shows the gonozooid of SMF 25285 (Voigt collection number 9797).

Fig. 2
Click on thumbnail to enlarge the image.

Figure 2: a-b Crisidia lagaaiji Voigt, 1987, holotype, SMF 25533, Danian, Ciply in the municipality Mons, Wallonia, Belgium. c-d Polyascosoecia pseudolichenoides Voigt, 1987, SMF 25553, Danian, Ciply in the municipality Mons, Wallonia. e-f Reteporidea georgiknetschi Voigt, 1984, holotype, SMF 25297, early Maastrichtian (Belemnitella junior belemnite Zone), Blom Quarry near Terblijt in the municipality Valkenburg aan de Geul, Limburg, Netherlands.
Scale bars: e 1 mm; c-d 500 µm; f 250 µm; a-b 100 µm.

Family Cytididae Orbigny, 1854

Genus Bicavea Orbigny, 1853

'Bicavea' fungiformis (Hagenow, 1846)

(Fig. 3a–b )

*p# 1846 Ceriopora fungiformis v. Hag.Hagenow, p. 595 (non Pl. XXIII.b, fig. 9).

# 1909 Ceriopora fungiformis (von Hagenow), 1846Gregory, p. 259

# 1959a Bicavea? fungiformis (v. Hagenow), 1846Voigt, p. 51, Pl. III, figs. 2–3.

p# 1972 Discocytis infundibuliformis (Hennig) – Brood, p. 345 (non Pl. XXXIV, figs. 6–7, Pl. XXXV, figs. 3, 4, 6).

# 2000 Bicavea fungiformis (Hagenow, 1846) – Viskova, p. 29, Pl. III, figs. 2–7.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany; Balsberg north of Kristianstad, Skåne län, Sweden.

Stratum typicum: Early Campanian and early Maastrichtian.

Neotype: SMF 26394 (Voigt, 1959a, Pl. III, figs. 2–3).

Original label: Voigt collection number 391.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Further distribution: Middle Campanian, quarry near Hrodna/Grodno (Гродна/Гродно), Grodno Region, Belarus.

Stratigraphical range: Middle Campanian to early Maastrichtian.

Remarks: The original imaged by Hagenow (1846), which probably came from Scania in Sweden and not from Rügen in Germany, could be conspecific with Lichenopora infundibuliformis Hennig, 1894, in which case 'Bicavea' fungiformis would be a senior synonym of L. infundibuliformis. Voigt (1959a) therefore suggested keeping the name L. infundibuliformis for specimens from Scania and restricting 'B.' fungiformis to specimens from Rügen.

Genus Cytidmonea Voigt, 1989

Cytidmonea kaeveri Voigt, 1989

(Fig. 3c–d )

*# 1989a Cytidmonea kaeveri n.g. n.sp. – Voigt, p. 93, Pl. 3, figs. 1–9, Pl. 4, figs. 1–6.

Holotype: Not found (Voigt, 1989a, Pl. 3, fig. 1).

Original label: Voigt collection number 11921.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: Cytidmonea kaeveri is the type species of Cytidmonea Voigt, 1989. The holotype was listed by Eiserhardt (1998).

Genus Discocytis Orbigny, 1854

Discocytis bettenstaedti Voigt, 1951

(Fig. 3e–f )

?# 1894 Lichenopora infundibuliformis n.sp. – Hennig, p. 35, Pl. 2, figs. 31–32.

?# 1909 Lichenopora infundibuliformis Hennig, 1894Gregory, p. 259.

*# 1951 Discocytis bettenstaedti n.sp. – Voigt, p. 41, Pl. 3, figs. 6–9.

?# 1972 Discocytis infundibuliformis (Hennig) – Brood, p. 345, Pl. XXXIV, figs. 6–7, Pl. XXXV, figs. 3, 4, 6.

Holotype: SMF 26304 (Voigt, 1951, Pl. 3, fig. 6).

Original label: Voigt collection number 255.

Locus typicus: Sehnde-Ilten, Lower Saxony, Germany.

Stratum typicum: Bryozoan-rich rubbly limestone in the Belemnitella junior belemnite Zone.

Further distribution: Late Maastrichtian, Saint-Symphorien and borehole near Jemappes (both Mons municipality, Wallonia, Belgium); Voerendaal-Kunrade, Limburg, Netherlands.

Stratigraphical range: Late Maastrichtian.

Remarks: Discocytis bettenstaedti was reported in the late Maastrichtian chalk tuff of Belgium and the Netherlands by Voigt (1957a). It is possibly a junior synonym of D. infundibuliformis (Hennig, 1894) from the late Campanian of Scania, Sweden, according to Brood (1972).

Fig. 3
Click on thumbnail to enlarge the image.

Figure 3: a-b 'Bicavea' fungiformis (Hagenow, 1846), neotype, SMF 26394, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Cytidmonea kaeveri Voigt, 1989, SMF 25783, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Discocytis bettenstaedti Voigt, 1951, holotype, SMF 26304, late Maastrichtian (Belemnitella junior belemnite Zone), Sehnde-Ilten, Lower Saxony, Germany.
Scale bars: a, c 1 mm; b, d-e 500 µm; f 100 µm.

Genus Discotruncatulipora Flor, 1975

Discotruncatulipora corbis Flor, 1975

(Fig. 4a–b )

*# 1975 Discotruncatulipora corbis n.sp. – Flor, p. 103, Pl. 7, figs. 1–6, Pl. 8, figs. 1–5.

Holotype: SMF 26460 (Flor, 1975, Pl. 8, fig. 1).

Original label: Voigt collection number 7653.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Further distribution: Cenomanian, Kaňk near Kutná Hora and further localities, Central Bohemian Region, Czech Republic. Late Cenomanian, Saint-Calais, Sarthe, Pays de la Loire, France. Early Turonian, quarry north of Předboj, Central Bohemian Region, Czech Republic; Hoher Stein in Dresden-Plauen, Saxony, Germany. Late Maastrichtian, Voerendaal-Kunrade, Limburg, Netherlands.

Stratigraphical range: Early Cenomanian to Late Maastrichtian.

Remarks: Discotruncatulipora corbis is the type species of Discotruncatulipora Flor, 1975. Flor (1975, p. 101) cited the Voigt collection number 7651 (SMF 26461), which is depicted in Pl. 7, figs. 4–6, as the "generotype" of Discotruncatulipora, but a different specimen – Voigt collection number 7653 (SMF 26460), which is depicted in Pl. 8, fig. 1 – as the holotype of D. corbis. Such a selection of a specimen as the "generotype" but of another specimen as the holotype of the type species is not considered in the International Code of Zoological Nomenclature (ICZN, 1964). Hence, the holotype of D. corbis is deemed to be SMF 26460 and the "generotype" SMF 26461 is considered to have no nomenclatural significance.

Heterocrisina candelabrum Voigt, 1987

(Fig. 4c–d )

*# 1987b Heterocrisina candelabrum n.sp. – Voigt, p. 42, Fig. 1A–H.

Holotype: SMF 25495 (Voigt, 1987b, Fig. 1A–B).

Original label: Voigt collection number 8610.

Locus typicus: Sint-Pietersberg near Maastricht, Limburg, Netherlands.

Stratum typicum: Meerssen Limestone IVf (Belemnitella junior belemnite Zone).

Further distribution: Late Maastrichtian, Curfs Quarry near Berg and Blom Quarry near Terblijt (both Valkenburg aan de Geul municipality); ENCI (Eerste Nederlandse Cement Industrie) pit on the hill Sint-Pietersberg (all Limburg, Netherlands).

Stratigraphical range: Late Maastrichtian.

Remarks: In the original description, the specimen labelled Voigt collection number 4564 (SMF 25490) is indicated as the holotype. However, the figure captions and the original label prove that Voigt collection number 8610 (SMF 25495) is the holotype of Heterocrisina candelabrum. SMF 25490 is depicted as Fig. 1D in Voigt (1987b).

Genus Infundibulipora Brood, 1972

Infundibulipora huckriedei Voigt, 1977

(Fig. 4e–f )

*# 1977b Infundibulipora huckriedei n.sp. – Voigt, p. 231, Pl. I, figs. 1–6, Pl. II, figs. 1–7, Pl. III, figs. 1–4.

Holotype: SMF 24033 (Voigt, 1977b, Pl. I, figs. 1–2).

Original label: Voigt collection number 7387.

Stratum typicum: Green marl clays between limestone of Campanian or Maastrichtian age.

Locus typicus: Southern side of the valley of "Kischk" and "Kemunu", northwest of Kemunu, Darmanu Mountains, northwest of Kerman, Kerman Province, Iran.

Further distribution: Late Campanian (Belemnitella mucronata belemnite Zone), Hemmingslycke near Kristianstad; Staversvad near Kristianstad-Arkelstorp (both Skåne län, Sweden). Late Maastrichtian, Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands. Late Maastrichtian (Belemnitella junior belemnite Zone), Voerendaal-Kunrade, Limburg, Netherlands.

Stratigraphical range: Late Campanian to late Maastrichtian.

Remarks: Prof. Dr R. Huckriede collected the holotype and donated it to Voigt.

Fig. 4
Click on thumbnail to enlarge the image.

Figure 4: a-b Discotruncatulipora corbis Flor, 1975, holotype, SMF 26460, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. c-d Heterocrisina candelabrum Voigt, 1987, holotype, SMF 25495, late Maastrichtian (Belemnitella junior belemnite Zone), Sint-Pietersberg near Maastricht, Limburg, Netherlands. e-f Infundibulipora huckriedei Voigt, 1977, holotype, SMF 24033, Campanian or Maastrichtian, southern side of the valley of "Kischk" and "Kemunu" northwest of the village Kemunu in the Darmanu Mountains northwest of Kerman, Kerman Province, Iran.
Scale bars: a, c, e 1 mm; f 500 µm; b, d 250 µm.

Genus Osculipora Orbigny, 1849

Osculipora iltenensis (Voigt, 1951)

(Fig. 5a–b )

*# 1951 Desmepora iltenensis n.sp. – Voigt, p. 41, Pl. 5, figs. 1–7.

?# 1972 Osculipora iltensis (Voigt), 1951Brood, p. 344, Pl. XXXIV, figs. 1–5, Pl. XXXV, fig. 1

Holotype: SMF 26385 (Voigt, 1951, Pl. 5, figs. 1–3).

Original label: Voigt collection number 275.

Locus typicus: Sehnde-Ilten, Lower Saxony, Germany.

Stratum typicum: Bryozoan-rich Rubbly limestone in the Belemnitella junior belemnite Zone.

Further distribution: According to Brood (1972, p. 344f): Santonian, several localities in southern Sweden. Early Campanian, Balsberg north of Kristianstad, Skåne län, Sweden. Late Campanian, Malen, Båstad Municipality, Skåne län, Sweden.

Stratigraphical range: Campanian to Maastrichtian.

Remarks: Brood (1972) reported more than 25 specimens of Osculipora iltensis from Campanian and Santonian sediments in Sweden but left open the possibility that the Campanian species could represent a new species because the autozooids are arranged irregularly between the alternating fascicles or in bundles.

Osculipora prescheri Voigt, 1998

(Fig. 5c–d )

# 1886 Truncatula truncata, GoldfussPergens & Meunier, p. 222.

# 1890 Osculipora truncata Goldfuss sp. – Osswald, p. 105.

p# 1925 Osculipora truncata Goldf.Levinsen, p. 439.

p# 1972 Osculipora truncata (Goldfuss), 1827Brood, p. 340, Pl. XLVIII, figs. 1–4 (non Pl. XLIV, figs. 11–12, Pl. XLVIII, figs. 5–7, Pl. LII, fig. 1).

*# 1998 Osculipora prescheri n.sp. – Voigt, p. 294, Fig. 2, Pl. 1, figs. 1–3, Pl. 2, figs. 1–4.

Holotype: SMF 26204 (Voigt, 1998, Pl. 1, figs. 1–3).

Original label: Voigt collection number 14481.

Locus typicus: Klintholm, island of Funen, South Denmark Region, Denmark.

Stratum typicum: Bryozoan limestone of late Danian age.

Further distribution: Late Danian, Faxe quarries, Zealand Region, Denmark; locality called "Ny/Klostergard/Jütland" in Denmark; glacial drift deposits on the island of Fehmarn, Schleswig-Holstein, Germany; Limhamn, Skåne län, Sweden.

Stratigraphical range: Danian.

Remarks: All specimens described in the literature as Osculipora truncata (Goldfuß, 1826) from the Danian were considered by Voigt (1998) to belong to O. prescheri.

Osculipora subhercynica (Voigt, 1924)

(Fig. 5e–f )

non# 1840 Idmonea semicylindrica N. – Roemer, Pl. V, fig. 21.

# 1841 Idmonea semicylindrica N. – Roemer, p. 20.

*# 1924a Desmepora subhercynica n.sp. – Voigt, p. 150, Pl. III, figs. 10–12.

# 1973a Desmepora subhercynica Voigt, 1924 Voigt, p. 124, p. 9, Figs. 1–2.

Lectotype (defined in Voigt, 1973a): the specimen he had depicted in 1924 (Voigt, 1924a, Pl. III, figs. 10–11). This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Neotype: SMF 24445 (Voigt, 1973a, Pl. 9, figs. 1–2).

Original label: Voigt collection number 6710.

Locus neotypicus: Burgberg west of Gehrden, Lower Saxony, Germany.

Stratum neotypicum: Late Santonian.

Further distribution: Santonian, abandoned iron-ore mine near Vechelde-Vallstedt; Barbara pit near Lengede-Barbecke; Lengede-Broistedt; Söhlde-Hoheneggelsen; (Ober-)Emscher Formation near Goslar-Oker; Sudmerberg near Goslar; Ilsede-Groß Bülten (all Lower Saxony, Germany). Early Campanian (Gonioteuthis quadrata belemnite Zone), Blankenburg (Harz); Wernigerode-Benzingerode (both Saxony-Anhalt, Germany).

Stratigraphical range: Middle Santonian to early Campanian.

Remarks: W. Pockrandt collected the neotype and gave it to Prof. Dr E. Voigt. The specimens of Idmonea semicylindrica Roemer, 1840, from Gehrden were referred to O. subhercynica (Voigt, 1924) and differ from the specimens from Rügen described in Voigt (1973a). The genus Desmepora Lonsdale, 1850, has been regarded as a junior synonym Osculipora Orbigny, 1849, by Brood (1972) and Taylor and McKinney (2006).

Fig. 5
Click on thumbnail to enlarge the image.

Figure 5: a-b Osculipora iltenensis (Voigt, 1951), holotype, SMF 26385, late Maastrichtian (Belemnitella junior belemnite Zone), Sehnde-Ilten, Lower Saxony, Germany. c-d Osculipora prescheri Voigt, 1998, holotype, SMF 26204, late Danian, Klintholm on the island of Funen, South Denmark Region, Denmark. e-f Osculipora subhercynica (Voigt, 1924), neotype, SMF 24445, late Santonian, Burgberg west of Gehrden, Lower Saxony, Germany.
Scale bars: c, e 1 mm; a, f 500 µm; d 250 µm; b 100 µm.

Family Petaloporidae Gregory, 1899

Genus Cavarinella Marsson, 1887

Cavarinella ramosa Marsson, 1887

(Fig. 6a–b )

non# 1851 Cavaria ramosa, Hag.Hagenow, p. 53, Pl. VI, fig. 1.

non# 1881 Cavaria ramosa v. Hag.Hamm, p. 25.

non# 1881 Cavaria ramosa Hag.Quenstedt, p. 276.

*# 1887 Cavarinella ramosa v. Hagenow sp. – Marsson, p. 19, Pl. I, fig. 6.

non# 1899 Cavaria ramosa, von Hagenow, 1851Gregory, p. 399, Fig. 54.

# 1922 Cavarinella ramosa Marsson, 1887Canu & Bassler, p. 135, Pl. 19, fig. 10.

p# 1925 Heteropora reticulata MarssonLevinsen, p. 428.

# 1935 Cavarinella ramosa Hagenow, 1851Bassler, p. 66.

# 1953 Cavarinella ramosa Hag., 1851Bassler, p. G63, Fig. 30.2.

# 1972 Reptomulticlausa ramosa (Hagenow), 1887Brood, p. 207, Pl. XXI, figs. 5–8, Pl. LXII, fig. 9.

# 1982a Cavarinella ramosa sensu Marsson, 1887 (non v. Hagenow, 1851) – Voigt, p. 73, Pl. 14, fig. 11, Pl. 15, figs. 1–7, Pl. 20, figs. 5–7.

Holotype: The material of Marsson (1887) belonged to the collections of the Preußische Geologische Landesanstalt (Prussian Geological Land Survey) that Voigt (1982a) reported to have been destroyed during World War II. However, part of the collection has been recovered (Martha, 2014). Should the type material of C. ramosa exist among the recovered material, the neotype would have to be set aside.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 24929 (Voigt, 1982a, Pl. 14, figs. 8–10).

Original label: Voigt collection number 9268.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Further distribution: Campanian, Lägerdorf, Schleswig-Holstein, Germany. Early Maastrichtian, Island of Møn, Zealand Region, Denmark; Island of Rügen, Mecklenburg-Vorpommern, Germany. Late Maastrichtian, Stevns Klint in the Stevns Kommune, Zealand Region, Denmark; Hemmoor, Lower Saxony, Germany and very common in glacial drift deposits from northern Germany containing white chalk or flint; Kvarnby, city district Husie of Malmö, Skåne län, Sweden. Danian, very common in white chalk exposures and in glacial drift deposits containing white chalk of Danian age in Denmark, Gemany and Sweden.

Stratigraphical range: Early Campanian to late Danian.

Remarks: Cavarinella ramosa Marsson, 1887 is the type species of Cavarinella Marsson, 1887. Canu and Bassler (1922) recognized that when Marsson (1887) chose Cavaria ramosa Hagenow, 1851, as the type species of his new genus Cavarinella, he described a completely different species than the Hagenow species. However, in his Fossilium Catalogus (Bassler, 1935) and in the Treatise (Bassler, 1953), Bassler gave C. ramosa Hagenow, 1851, as the type species of Cavarinella. The case was clarified by Voigt (1982a). The true Cavaria ramosa Hagenow, 1851, was regarded as a subjective synonym of Lopholepis radians Hagenow, 1851. To maintain the genus Cavarinella, Voigt (1982a) chose C. ramosa sensu Marsson (1887) from the white chalk of Rügen as the type species.

Genus Laterocavea Orbigny, 1853

Laterocavea distorta Voigt, 1989

(Fig. 6c–d )

*# 1989a Laterocavea distorta n.sp. – Voigt, p. 95, Pl. 5, figs. 1–8, Pl. 6, figs. 1–2.

# 1995a Laterocavea distorta VoigtVoigt, unnumbered figure, sample top left.

Holotype: SMF 25809 (Voigt, 1989a, Pl. 5, figs. 3–6).

Original label: Voigt collection number 11866.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation, in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Neostlingoceras carcitanense ammonite Zone of the earliest Cenomanian.

Fig. 6
Click on thumbnail to enlarge the image.

Figure 6: a-b Cavarinella ramosa Marsson, 1887, neotype, SMF 24929, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Laterocavea distorta Voigt, 1989, holotype, SMF 25809, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: c 1 mm; a, d 500 µm; b 100 µm.

Suborder Cerioporina Hagenow, 1851

Family Cerioporidae Busk, 1859

Genus Cryptoglena Marsson, 1887

Cryptoglena adspersa Marsson, 1887

(Fig. 7a–b )

*# 1887 Cryptoglena adspersa n.sp. – Marsson, p. 16, Pl. I, fig. 3.

# 1899 Cryptoglena adspersa, Marsson, 1887Gregory, p. 405.

# 1935 Cryptoglena adspersa Marsson, 1887Bassler, p. 85.

# 1953 Cryptoglena adspersa Marsson, 1887Bassler, p. G75, Fig. 39.3.

# 1982a Cryptoglena adspersa Marsson 1887Voigt, p. 78, Pl. 17, figs. 1–9, Pl. 19, fig. 5.

Holotype: The material of Marsson (1887) belonged to the collections of the Preußische Geologische Landesanstalt (Prussian Geological Land Survey) that Voigt (1982a) reported to have been destroyed during World War II. However, part of the collection has been recovered (Martha, 2014), and if the type material for C. adspersa belongs to the recovered material, the neotype would have to be set aside.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 24920 (Voigt, 1982a, Pl. 17, fig. 1).

Original label: Voigt collection number 9260.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Further distribution: Late Campanian, Meudon, Île-de-France, France. Early Maastrichtian, Island of Møn, Zealand Region, Denmark; Island of Rügen, Mecklenburg-Vorpommern, Germany. Late Maastrichtian, Hemmoor, Lower Saxony, Germany; rare in glacial drifts containing white chalk in northern Germany; Tuffeau de Maastricht, Maastricht area, Limburg, Netherlands. Danian, glacial drifts near Neu Wulmstorf, Lower Saxony, Germany. Late Middle Danian, Voldum, Favrskov Kommune, Region Midtjylland, Denmark.

Stratigraphical range: Late Campanian to Danian.

Remarks: Cryptoglena adspersa is the type species of Cryptoglena Marsson, 1887.

Genus Marssoniella Levinsen, 1925

Marssoniella cenomana Voigt, 1974

(Fig. 7c–d )

*# 1974 Marssoniella cenomana n.sp. – Voigt, p. 202, Figs. 1–3, 5–8.

# 1995a Marssoniella cenomana VoigtVoigt, unnumbered figure, samples bottom left and bottom right.

Holotype: SMF 24549 (Voigt, 1974, Figs. 1–2).

Original label: Voigt collection number 6661.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation, in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: B.A. Engel collected the holotype and gave it to Voigt.

Genus Pachyteichopora Levinsen, 1925

Pachyteichopora bugei Voigt, 1978

(Fig. 7e–f )

*# 1978 Pachyteichopora bugei n.g. n.sp. – Voigt, p. 265, Figs. 14–20.

# 1984 Pachyteichopora bugei Voigt, 1978Boardman, Fig. 10D–F.

# 1985a Pachyteichopora bugei Voigt, 1978Voigt, p. 629, Pl. 2, figs. 3–4.

Holotype: SMF 24690 (Voigt, 1978, Fig. 20).

Original label: Voigt collection number 8466.

Locus typicus: See remarks.

Stratum typicum: Coniacian.

Further distribution: Coniacian, Saint-Christophe-sur-le-Nais, Indre-et-Loire; Tours, Indre-et-Loire (both Centre-Val de Loire, France); Fécamp, Normandy, France. Coniacian to Santonian, Gorges du Nant near Cognin-les-Gorges, Isère, Auvergne-Rhône-Alpes, France; Saint-Paterne-Racan, Indre-et-Loire, Centre-Val de Loire, France.

Stratigraphical range: Coniacian to Campanian.

Remarks: Villedieu (Indre et Loire) is given as the provenance by Voigt (1978) in his description. However, the original label indicates that the holotype comes from Tours (Indre-et-Loire). Pachyteichopora bugei was reported from the Coniacian (to Santonian) of Saint-Paterne-Racan and also occurs in the Campanian of the Aquitaine Basin according to Voigt (1985a).

Fig. 7
Click on thumbnail to enlarge the image.

Figure 7: a-b Cryptoglena adspersa (Marsson, 1887), neotype, SMF 24920, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Marssoniella cenomana Voigt, 1974, holotype, SMF 24549, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Pachyteichopora bugei Voigt, 1978, holotype, SMF 24690, Coniacian, probably Tours, Centre-Val de Loire, France.
Scale bars: c 2.5 mm; e 1 mm; a, d 500 µm; f 250 µm; b 100 µm.

Pachyteichopora punctata Voigt, 1978

(Fig. 8a–b )

*# 1978 Pachyteichopora punctata n.g. n.sp. – Voigt, p. 267, Figs. 22–28.

Holotype: SMF 24683 (Voigt, 1978, Figs. 22–23).

Original label: Voigt collection number 8467.

Locus typicus: Båstad, Skåne län, Sweden.

Stratum typicum: Rubbly limestone from the Belemnitella mucronata belemnite Zone.

Stratigraphical range: Late Campanian.

Genus Stylodefranciopora Voigt & Vávra, 2006

Stylodefranciopora turris Voigt & Vávra, 2006

(Fig. 8c–d )

*# 2006 Stylodefranciopora turris nov. gen. nov. spec. – Voigt & Vávra, p. 138, Pl. 1, figs. 1–8, Pl. 2, figs. 1–7, Pl. 3, figs. 1–4.

Holotype: SMF 26248 (Voigt & Vávra, 2006, Pl. 1, fig. 1).

Original label: Voigt collection number 10326 (SMF 3009).

Locus typicus: Saturn pit near Kronsmoor, Schleswig-Holstein, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Paratypes: SMF 26249–26256.

Further distribution: Santonian, Burgberg west of Gehrden, Lower Saxony, Germany. Campanian, chalk pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany. Early Maastrichtian, Island of Møn, Zealand Region, Denmark.

Stratigraphical range: Santonian to early Maastrichtian.

Family Corymboporidae Smitt, 1867

Genus Amphimarssoniella Voigt, 1974

Amphimarssoniella klaumanni Voigt, 1974

(Fig. 8e–f )

*# 1974 Amphimarssoniella klaumanni n.g. n.sp. – Voigt, p. 209, Figs. 9–28.

# 1981a Amphimarssoniella klaumanni VoigtVoigt, Fig. 3a–c.

# 1995a Amphimarssoniella klaumanni VoigtVoigt, unnumbered figure, sample top right.

# 1999 Amphimarssoniella klaumanni Voigt, 1974Taylor & Grischenko, Fig. 18.

Holotype: SMF 24533 (Voigt, 1974, Fig. 11).

Original label: Voigt collection number 6665.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation, in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: Amphimarssoniella klaumanni is the type species of Amphimarssoniella Voigt, 1974. H. Klaumann collected the holotype and gave it to Voigt.

Fig. 8
Click on thumbnail to enlarge the image.

Figure 8: a-b Pachyteichopora punctata Voigt, 1978, holotype, SMF 24683, late Campanian (Belemnitella mucronata belemnite Zone), Båstad, Skåne län, Sweden. c-d Stylodefranciopora turris Voigt & Vávra, 2006, holotype, SMF 26248, early Maastrichtian, Saturn pit near Kronsmoor, Schleswig-Holstein, Germany. e-f Amphimarssoniella klaumanni Voigt, 1974, holotype, SMF 24533, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: a, e 2.5 mm; c 1 mm; f 500 µm; d 250 µm; b 100 µm.

Suborder Melicerititida Pergens, 1889

Family Eleidae Orbigny, 1853

Genus Biforicula Voigt, 1989

Biforicula acuminata (Voigt, 1973)

(Fig. 9a–b )

p# 1924a Elea nodulifera n.sp. – Voigt, p. 165, Pl. IV, fig. 8 (non figs. 7, 9).

*# 1973a Foricula acuminata n.sp. – Voigt, p. 139, Pl. 14, figs. 5–7.

Holotype: SMF 26436 (Voigt, 1973a, Pl. 14, fig. 7).

Original label: Voigt collection number 6674a.

Locus typicus: Abandoned Barbara pit near Lengede-Barbecke, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Further distribution: Santonian, hills Sudmerberg near Goslar; Burgberg west of Gehrden; abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten; abandoned iron-ore mine near Vechelde-Vallstedt; Lengede-Broistedt; Söhlde-Hoheneggelsen; Harzburg; Bad Harzburg-Harlingerode (all Lower Saxony, Germany).

Stratigraphical range: Middle Santonian.

Remarks: In both the German and the English versions of the the abstract of Voigt (1973a), the name of this species is incorrectly given as "Foricula acutimargo n.sp.", which is a nomen nudum.

Biforicula? nodulifera (Voigt, 1924)

(Fig. 9c–d )

p# 1924a Elea nodulifera n.sp. – Voigt, p. 165, Pl. IV, figs. 7, 9 (non fig. 8).

*# 1973a Foricula nodulifera (Voigt, 1924) – Voigt, p. 138, Pl. 14, figs. 1–4.

Lectotype (defined in Voigt, 1973a): the specimen figured by Voigt (1924a, Pl. IV, fig. 7). This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Sudmerberg near Goslar, Lower Saxony, Germany.

Stratum typicum: Sponge-rich marls of the Emscher Formation, early Santonian.

Neotype: SMF 26436 (Voigt, 1973a, Pl. 14, fig. 1).

Original label: Voigt collection number 6761 (= Voigt collection number 6760).

Locus neotypicus: Burgberg west of Gehrden, Lower Saxony, Germany.

Stratum neotypicum: Late Santonian.

Further distribution: Middle Santonian, Salzberg near Quedlinburg; abandoned Barbara pit near Lengede-Barbecke; abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten; Bad Harzburg; abandoned iron-ore mine near Vechelde-Vallstedt; Lengede-Broistedt; Söhlde-Hoheneggelsen (all Lower Saxony, Germany).

Stratigraphical range: Santonian.

Remarks: The label with the neotype has the Voigt collection number 6760.

Genus Elea Orbigny, 1853

Elea elegantula Taylor, 1994

(Fig. 9e–f )

*# 1994 Elea elegantula sp. nov. – Taylor, p. 19, Figs. 29–43.

Holotype: SMF 26499 (Taylor, 1994, Figs. 29, 36, 39–42).

Original label: Voigt collection number 10451.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Paratypes: SMF 26499 (in same cavity slide as holotype), 26500.

Stratigraphical range: Earliest Cenomanian.

Remarks: The paratypes with the Voigt collection numbers 9847 and 9848 were not found.

Fig. 9
Click on thumbnail to enlarge the image.

Figure 9: a-b Biforicula acuminata (Voigt, 1973), holotype, SMF 26436, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned Barbara pit near Lengede-Barbecke, Lower Saxony, Germany. c-d Biforicula ? nodulifera (Voigt, 1924), neotype, SMF 26436, late Santonian, Burgberg west of Gehrden, Lower Saxony, Germany. e-f Elea elegantula Taylor, 1994, holotype, SMF 26499, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: a, c, e 1 mm; b 250 µm; d, f 100 µm.

Elea flabellata Taylor, 1994

(Fig. 10a–b )

*# 1994 Elea flabellata sp. nov. – Taylor, p. 20, Figs. 44–48.

Holotype: SMF 26501 (Taylor, 1994, Figs. 44–48).

Original label: Voigt collection number 10448.

Locus typicus: Abandoned iron ore pit Mathilde near Lengede-Broistedt, Lower Saxony, Germany.

Stratum typicum: Early Santonian.

Stratigraphical range: Early Santonian.

Remarks: The holotype is the only reported specimen of this species.

Elea mackinneyi Taylor, 1994

(Fig. 10c–d )

*# 1994 Elea mackinneyi sp. nov. – Taylor, p. 26, Figs. 66–71.

Holotype: SMF 26503 (Taylor, 1994, Figs. 66–71).

Original label: Voigt collection number 10474.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Paratype: SMF 26504.

Stratigraphical range: Earliest Cenomanian.

Elea pseudolamellosa Taylor, 1994

(Fig. 10e–f )

*# 1994 Elea pseudolamellosa sp. nov. – Taylor, p. 30, Figs. 72–78.

Holotype: SMF 26506 (Taylor, 1994, Figs. 75, 78).

Original label: Voigt collection number 10462.

Locus typicus: "St Severin d'Uzel", probably Chenac-Saint-Seurin-d'Uzet, Charente-Maritime, Nouvelle-Aquitaine, France.

Stratum typicum: Late Campanian.

Paratypes SMF 26505, 26507.

Further distribution: Campanian, railway station in Soullans, Vendée, Pays de la Loire, France. Late Campanian, Le Cailleau near Talmont-sur-Gironde, Charente-Maritime, Nouvelle-Aquitaine, France.

Stratigraphical range: Campanian.

Remarks: Taylor (1994) indicated "St Severin d'Uzel" as the type locality. However, this probably represents an incorrect transcription of the original handwritten label and the true locality is likely to be Saint-Seurin-d'Uzet, a part of the commune of Chenac-Saint-Seurin-d'Uzet.

Fig. 10
Click on thumbnail to enlarge the image.

Figure 10: a-b Elea flabellata Taylor, 1994, holotype, SMF 26501, early Santonian, abandoned iron ore pit Mathilde near Lengede-Broistedt, Lower Saxony, Germany. c-d Elea mackinneyi Taylor, 1994, holotype, SMF 26503, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Elea pseudolamellosa Taylor, 1994, holotype, SMF 26506, late Campanian, "St Severin d'Uzel" (?), Nouvelle-Aquitaine, France.
Scale bars: a 2.5 mm; c 1 mm; e 500 µm; d 250 µm; b, f 100 µm.

Elea viskovae Taylor, 1994

(Fig. 11a–b )

*# 1994 Elea viskovae sp. nov. – Taylor, p. 38, Figs. 95–99.

Holotype: SMF 26508 (Taylor, 1994, Figs. 95–99).

Original label: Voigt collection number 10450.

Locus typicus: A locality called "Kyzylsaj", ca. 180 km east of Fort-Shevchenko (Форт-Шевченко), Mengystau Province, Kazakhstan.

Stratum typicum: Turonian.

Paratype: SMF 26509.

Stratigraphical range: Turonian.

Elea whiteleyi Taylor, 1994

(Fig. 11c–d )

*# 1994 Elea whiteleyi sp. nov. – Taylor, p. 38, Figs. 100–109.

Holotype: SMF 26511 (Taylor, 1994, Figs. 102–103).

Original label: Voigt collection number 10538.

Locus typicus: Saint-Germain-la-Campagne, Normandy, France.

Stratum typicum: Base of the middle Cenomanian.

Paratypes: SMF 26510, 26512, 26513

Further distribution: Early (?) Cenomanian, Saint-Jouin-Bruneval, Seine-Maritime, Normandy, France.

Stratigraphical range: Early to middle Cenomanian.

Genus Meliceritella Levinsen, 1925

Meliceritella schneemilchae Taylor, 1987

(Fig. 11e–f )

*# 1987 Meliceritella schneemilchae sp. nov. – Taylor, p. 72, Pl. 1a–i.

Holotype: SMF 26495 (Taylor, 1987, Pl. 1c–e).

Original label: Voigt collection number 10328.

Locus typicus: Schinkel pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.

Stratum typicum: Offaster pilula echinoid Zone in the early Campanian.

Paratypes: SMF 26493, 26494, 26496–26498.

Further distribution: Early Campanian, Alemannia quarry near Sehnde-Höver, Lower Saxony, Germany.

Stratigraphical range: Early Campanian.

Fig. 11
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Figure 11: a-b Elea viskovae Taylor, 1994, holotype, SMF 26508, Turonian, locality called "Kyzylsaj", ca. 180 km east of Fort-Shevchenko (Форт-Шевченко), Mengystau Province, Kazakhstan. c-d Elea whiteleyi Taylor, 1994, holotype, SMF 26511, middle Cenomanian, Saint-Germain-la-Campagne, Normandy, France. e-f Meliceritella schneemilchae Taylor, 1987, holotype, SMF 26495, early Campanian (Offaster pilula echinoid Zone), Schinkel pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.
Scale bars: e 1 mm; a 500 µm; b-c 250 µm; f 100 µm; d 50 µm.

Genus Meliceritites Roemer, 1841

Meliceritites eleoides Voigt, 1975

(Fig. 12a–b )

*# 1975a Meliceritites eleoides n.sp. – Voigt, p. 243, Pl. 1, figs. 3–10.

Holotype: SMF 24578 (Voigt, 1975a, Pl. 1, figs. 8–10).

Original label: Voigt collection number 7428.

Locus typicus: Drilling in the chalk pit Schinkel-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.

Stratum typicum: White chalk of the Uintacrinus socialis crinoid Zone (late Santonian).

Further distribution: Early Campanian, Sehnde-Höver, Lower Saxony, Germany.

Stratigraphical range: Santonian to base of the late Campanian.

Remarks: The specimens assigned by Voigt (1975a) to Meliceritites eleoides are all small fragments that lack primary eleozooids and opercula. However, the straight proximal edges of the autozooidal apertures, and the occurrence of a possible secondary nanozooid (Fig. 64 b, upper right) support the assignment of this species to Meliceritites Roemer, 1841.

Meliceritites pseudoangulosa Voigt, 1973

(Fig. 12c–d )

# 1924a Meliceritites angulosa OrbignyVoigt, p. 168, Pl. IV, fig. 10.

*# 1973a Meliceritites pseudoangulosa n.sp. – Voigt, p. 135, Pl. 11, fig. 5, Pl. 12, figs. 5–9.

Holotype: SMF 26435 (Voigt, 1973a, Pl. 11, fig. 5).

Original label: Voigt collection number 6685.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Further distribution: Santonian, Ilsede-Groß Bülten; Gehrden; Sudmerberg near Goslar; abandoned iron-ore mine near Vechelde-Vallstedt; Lengede-Broistedt; Söhlde-Hoheneggelsen; Harzburg; Bad Harzburg-Harlingerode (all Lower Saxony, Germany).

Stratigraphical range: Santonian.

Genus Reptomultelea Orbigny, 1853

Reptomultelea acclivata Taylor, 1994

(Fig. 12e–f )

*# 1994 Reptomultelea acclivata sp. nov. – Taylor, p. 54, Figs. 148–154.

Holotype: SMF 26521 (Taylor, 1994, Figs. 148–151).

Original label: Voigt collection number 10427.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: The paratype bearing Voigt collection number 10502 was not found.

Fig. 12
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Figure 12: a-b Meliceritites ? eleoides Voigt, 1975, holotype, SMF 24578, late Santonian (Uintacrinus socialis crinoid Zone), Schinkel-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany. c-d Meliceritites pseudoangulosa Voigt, 1973, holotype, SMF 26435, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany. e-f Reptomultelea acclivata Taylor, 1994, holotype, SMF 26521, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: e 2.5 mm; c 1 mm; a 500 µm; d 250 µm, b, f 100 µm.

Reptomultelea convexa Taylor, 1994

(Fig. 13a–b )

*# 1994 Reptomultelea convexa sp. nov. – Taylor, p. 66, Figs. 191–196.

Holotype: SMF 26524 (Taylor, 1994, Figs. 191–193).

Original label: Voigt collection number 10467.

Locus typicus: Chenu, Pays de la Loire, France.

Stratum typicum: Turonian (?).

Paratypes: SMF 26525, 26526.

Further distribution: Turonian, Bois de Gareau near Écommoy, Pays de la Loire, France.

Stratigraphical range: Turonian.

Remarks: G. Breton collected the holotype and gave it to Voigt.

Reptomultelea parvula Taylor, 1994

(Fig. 13c–d )

*# 1994 Reptomultelea parvula sp. nov. – Taylor, p. 83, Figs. 243–248.

Holotype: SMF 26531 (Taylor, 1994, Figs. 243–248).

Original label: Voigt collection number 10434.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: H. Klaumann collected the holotype and gave it to Voigt. The holotype is the only reported specimen of this species.

Reptomultelea pegma Taylor, 1994

(Fig. 13e–f )

*# 1994 Reptomultelea pegma sp. nov. – Taylor, p. 85, Figs. 249–255.

Holotype: SMF 26532 (Taylor, 1994, Figs. 249–252, 255).

Original label: Voigt collection number 10430.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Paratypes: SMF 26532 (same cavity slide as holotype), 26533.

Stratigraphical range: Earliest Cenomanian.

Fig. 13
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Figure 13: a-b Reptomultelea convexa Taylor, 1994, holotype, SMF 26524, probably Turonian, Chenu, Pays de la Loire, France. c-d Reptomultelea parvula Taylor, 1994, holotype, SMF 26531, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Reptomultelea pegma Taylor, 1994, holotype, SMF 26532, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: a, c 1 mm; f 500 µm; b 250 µm; d-e 100 µm.

Reptomultelea pseudopalpebrosa Taylor, 1994

*# 1994 Reptomultelea pseudopalpebrosa sp. nov. – Taylor, p. 89, Figs. 263–268.

Holotype: Not found (Taylor, 1994, Figs. 263–268).

Original label: Voigt collection number 7057.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: The holotype is the only reported specimen of this species.

Reptomultelea reedi Taylor, 1994

(Fig. 14a–b )

*# 1994 Reptomultelea reedi sp. nov. – Taylor, p. 91, Figs. 269–274.

Holotype: SMF 26537 (Taylor, 1994, Figs. 269–274).

Original label: Voigt collection number 10429.

Locus typicus: Saint-Calais, Sarthe, Pays de la Loire, France.

Stratum typicum: Late Cenomanian.

Stratigraphical range: Late Cenomanian.

Remarks: The holotype is the only reported specimen of this species.

Reptomultelea scanica Taylor, 1994

(Fig. 14c–d )

*# 1994 Reptomultelea scanica sp. nov. – Taylor, p. 96, Figs. 287–296.

Holotype: SMF 26540 (Taylor, 1994, Figs. 287, 289–291, 295).

Original label: Voigt collection number 10441.

Locus typicus: Karlshamn, Blekinge län, Sweden.

Stratum typicum: Early Campanian (Belemnellocamax mammillatus belemnite Zone).

Paratypes: SMF 26541, 26542.

Stratigraphical range: Early Campanian.

Fig. 14
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Figure 14: a-b Reptomultelea reedi Taylor, 1994, holotype, SMF 26537, late Cenomanian, Saint-Calais, Sarthe, Pays de la Loire, France. c-d Reptomultelea scanica Taylor, 1994, holotype, SMF 26540, early Campanian (Belemnellocamax mammillatus belemnite Zone), Karlshamn, Blekinge län, Sweden.
Scale bars: a 1 mm; c 500 µm; d 250 µm; b 100 µm.

Suborder Rectangulata Waters, 1887

Family Lichenoporidae Smitt, 1867

Genus Doliocoitis Buge & Tillier, 1977

Doliocoitis atlantica Buge & Tillier, 1977

(Fig. 15a–b )

*# 1977 Doliocoitis atlantica n.sp. – Buge & Tillier, p. 4, Pl. I, figs. 1–6, Pl. II, figs. 1–5.

Holotype: SMF 26450 (Buge & Tillier, 1977, Pl. I, fig. 3).

Original label: № 6, Holotype (Voigt collection number 7498).

Locus typicus: Station 166 of the RV "Meteor" campaign 1973, southwest of the coast of Sierra Leone, 50 km west of Sherbro Island at 7°27'N, 13°37'W, depth 81 m.

Stratigraphical range: Recent.

Remarks: Doliocoitis atlantica is the type species of Doliocoitis Buge & Tillier, 1977.

Genus Lichenopora Defrance, 1823

Lichenopora costata (Hagenow, 1846)

(Fig. 15c–d )

*# 1846 Ceriopora costata v. Hag.Hagenow, p. 594, Pl. XXIII.b, fig. 8.

# 1959a Lichenopora costata (v. Hagenow), 1846Voigt, p. 50, Pl. III, fig. 1.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26393 (Voigt, 1959a, Pl. III, fig. 1).

Original label: Voigt collection number 390.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Lichenopora pedunculata Voigt, 1989

(Fig. 15e–f )

*# 1989a Lichenopora pedunculata n.sp. – Voigt, p. 96, Pl. 6, figs. 3–6, Pl. 7, figs. 1–8.

# 1992a Lichenopora pedunculata VoigtVoigt, Pl. 5, figs. 3–4.

# 1997 Lichenopora pedunculata Voigt, 1989Gordon & Taylor, Fig. 16.

Holotype: SMF 25810 (Voigt, 1989a, Pl. 7, fig. 3).

Original label: Voigt collection number 6835.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation, in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Fig. 15
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Figure 15: a-b Doliocoitis atlantica Buge & Tillier, 1977, holotype, SMF 26450, Recent, Station 166 of the RV "Meteor" campaign 1973 southwest of the coast of Sierra Leone, 50 km west of Sherbro Island at 7°27'N, 13°37'W at a depth of 81 m. c-d Lichenopora costata (Hagenow, 1846), holotype, SMF 26393, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. e-f Lichenopora pedunculata Voigt, 1989, holotype, SMF 25810, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: a, c 1 mm; d 500 µm; b, e-f 250 µm.

Suborder Tubuliporina Milne Edwards, 1838

Family Actinoporidae Vigneaux, 1949

Genus Actinopora Orbigny, 1853

Actinopora auei Voigt, 1996

(Fig. 16a–b )

*# 1996a Actinopora auei n.sp. – Voigt, p. 46, Pl. I, figs. 1–4, Pl. II, figs. 1–2, Pl. III, fig. 1.

Holotype: SMF 26144 (Voigt, 1996a, Pl. I, figs. 1–4, Pl. II, figs. 1–2, Pl. III, fig. 1).

Original label: Voigt collection number 14118.

Locus typicus: Scree on the beach near Ahrenshoop, Mecklenburg-Vorpommern, Germany.

Stratum typicum: Till of the Weichsel glaciation.

Stratigraphical range: Probably Campanian.

Remarks: J. Aue collected the holotype and gave it to. Voigt. The holotype, which is the only known specimen of this species, was found in a glacial drift deposit. Voigt (1996a) supposed the material to be derieved from the Swedish rubbly limestone often found inside glacial drift deposits in northern Germany, and therefore probably of Campanian age.

Family Celluliporidae Buge & Voigt, 1972

Genus Cellulipora Orbigny, 1849

Cellulipora westfalica Buge & Voigt, 1972

(Fig. 16c–d )

*# 1972 Cellulipora westfalica nov. sp. – Voigt, p. 141, Pl. 13, figs. 1–7.

# 1981a Cellulipora westfalica Buge & VoigtVoigt, Fig. 4a–c.

# 2004 Cellulipora westfalica Buge et VoigtViskova, Fig. 2.

# 2007 Cellulipora westfalica Buge et VoigtViskova, Pl. 8, figs. 3–4.

Holotype: SMF 24420 (Buge & Voigt, 1972, Pl. 13, fig. 1).

Original label: Voigt collection number 6031.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation, in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Family Diaperoeciidae Canu, 1918

Genus Desmediaperoecia Canu & Bassler, 1920

Desmediaperoecia europaea Voigt, 1987

(Fig. 16e–f )

*# 1987b Desmediaperoecia europaea n.sp. – Voigt, p. 52, Figs. 4A–D, 5A–D.

Holotype: SMF 25906 (Voigt, 1987b, Figs. 4A–D).

Original label: Voigt collection number 4609.

Locus typicus: Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands.

Stratum typicum: Belemnitella junior belemnite Zone, Tuffeau de Maastricht (Meerssen Limestone).

Further distribution: Late Maastrichtian, Sint-Pietersberg near Maastricht, Limburg, Netherlands.

Stratigraphical range: Late Maastrichtian.

Fig. 16
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Figure 16: a-b Actinopora auei Voigt, 1996, holotype, SMF 26144, probably Campanian, Ahrenshoop, Mecklenburg-Vorpommern, Germany. c-d Cellulipora westfalica Buge & Voigt, 1972, holotype, SMF 24420, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Desmediaperoecia europaea Voigt, 1987, holotype, SMF 25906, late Maastrichtian (Belemnitella junior belemnite Zone), Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands.
Scale bars: a 10 mm; b-c, e 1 mm; 250 µm; d 100 µm.

Genus Diaperoecia Canu, 1918

Diaperoecia incrustata (Hagenow, 1840)

(Fig. 17a–b )

*# 1840 Ceriopora incrustata Goldf.Hagenow, p. 647.

# 1909 Reptomulticava ? incrustata (von Hagenow), 1840Gregory, p. 148.

# 1959a Diaperoecia incrustata (v. Hagenow), 1840Voigt, p. 48, Pl. II, figs. 2–4.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26389 (Voigt, 1959a, Pl. II, figs. 2–3).

Original label: Voigt collection number 386.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Remarks: It is unclear why Hagenow assigned the authorship of his new species Ceriopora incrustata to Goldfuß since Goldfuß did not describe any species with this name. Subsequent authors have regarded Hagenow as the author of the species.

Diaperoecia neumeieri Voigt, 1995

(Fig. 17c–d )

*# 1995b Diaperoecia neumeieri n.sp. – Voigt, p. 11, Pl. 1, figs. 1–7, Pl. 2, figs. 1–4, Pl. 3, figs. 1–5, Pl. 4, figs. 1–4, Pl. 5, figs. 1–5.

Holotype: SMF 26050 (Voigt, 1995b, Pl. 2, fig. 4, Pl. 4, figs. 1–2).

Original label: Voigt collection number 13395.

Locus typicus: Schierling-Zaitzkofen, Bavaria, Germany.

Stratum typicum: Großberg Formation, in a field west of Zaitzkopf.

Stratigraphical range: Late Turonian.

Diaperoecia tubuliformis Voigt, 1973

(Fig. 17e–f )

# 1924a Microecia tubulus OrbignyVoigt, p. 136.

*# 1973a Diaperoecia tubuliformis (von Hagenow), 1840Voigt, p. 120, Pl. 8, figs. 4–7.

Holotype: SMF 24438 (Voigt, 1973a, Pl. 8, fig. 4).

Original label: Voigt collection number 6726.

Locus typicus: Abandoned iron-ore mine near Vechelde-Vallstedt, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Further distribution: Santonian, Burgberg west of Gehrden; Sudmerberg near Goslar; Barbara pit near Lengede-Barbecke; Lengede-Broistedt; abandoned iron-ore mine Groß-Bülten near Ilsede-Groß Bülten; Söhlde-Hoheneggelsen; Bad Harzburg (all Lower Saxony, Germany).

Stratigraphical range: Middle Santonian.

Remarks: Diastopora tubuliformis and D. tubulus Orbigny, 1851, differ only in the morphology of the gonozooid.

Fig. 17
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Figure 17: a-b Diaperoecia incrustata (Hagenow, 1840), neotype, SMF 26389, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Diaperoecia neumeieri Voigt, 1995, holotype, SMF 26050, late Turonian, Schierling-Zaitzkofen, Bavaria, Germany. e-f Diaperoecia tubuliformis Voigt, 1973, holotype, SMF 24438, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned iron-ore mine near Vechelde-Vallstedt, Lower Saxony, Germany.
Scale bars: e 5 mm; a 1 mm; c, f 500 µm; b, d 250 µm.

Family Entalophoridae Reuss, 1869

Genus Clavisparsa Orbigny, 1853

Clavisparsa huckei Voigt, 1962

(Fig. 18a–b )

*# 1962a Clavisparsa huckei n.sp.– Voigt, p. 250, Pl. 28, figs. 9–12.

# 2005 Clavisparsa huckei Voigt, 1962 – Hinz-Schallreuter & Schallreuter, p. 538.

Holotype: SMF 24141 (Voigt, 1962a, Pl. 28, fig. 9).

Original label: Voigt collection number 3542.

Locus typicus: Chalk in glacial drift deposit F near Neu Wulmstorf-Daerstorf, Lower Saxony, Germany.

Stratum typicum: Glacial drift deposit containing white chalk of late Maastrichtian age.

Further distribution: Late Maastrichtian, glacial drift F in a quarry near Tornesch, Schleswig-Holstein, Germany; glacial drift C containing flint near Neu Wulmstorf, Lower Saxony, Germany. Late Maastrichtian (Belemnella junior belemnite Zone), Hemmoor, Lower Saxony, Germany.

Stratigraphical range: Late Maastrichtian.

Genus Mecynoecia Canu, 1918

Mecynoecia (?) latedistans Voigt & Viaud, 1983

(Fig. 18c–d )

*# 1983 Mecynoecia (?) latedistans n.sp. – Voigt & Viaud, p. 222, Pl. 1, figs. 11–12.

Holotype: SMF 25488 (Voigt & Viaud, 1983, Pl. 1, fig. 11).

Original label: Voigt collection number 9823.

Locus typicus: Les Guignardières, Soullans, Vendée, Pays de la Loire, France.

Stratum typicum: Mid-late Santonian to early Campanian.

Further distribution: Coniacian, Joué-lès-Tours, Indre-et-Loire, Centre-Val de Loire, France. Santonian to Campanian, Le Glajou in Soullans, Vendée; Chemin de la Ruelle in Loissons, Saint-Hilaire-de-Riez, Vendée; Les Rosèliers in Notre-Dame-de-Riez, Vendée (all Pays de la Loire, France). Santonian, Vendôme, Loir-et- Cher, Centre-Val de Loire, France. Campanian, Railway station, Soullans, Vendée, Pays de la Loire, France.

Stratigraphical range: Coniacian to Campanian.

Remarks: The holotype is mounted on the same SEM stub as Voigt collection number 9910 (Pl. 1, fig. 12). The assignment of the species to the genus Mecynoecia Canu, 1918, is somewhat unclear since no specimens with gonozooids have yet been found.

Family Frondiporidae Busk, 1875

Genus Frondipora Link, 1807

Frondipora praecursor Voigt, 1951

(Fig. 18e–f )

*# 1951 Frondipora praecursor n.sp. – Voigt, p. 39, Pl. 4, figs. 18–20.

# 1972 Frondipora praecursor Voigt, 1953 [sic] – Brood, p. 250, Pl. XXVII, figs. 2–4.

Holotype: SMF 26323 (Voigt, 1951, Pl. 4, figs. 18–19).

Original label: Voigt collection number 273.

Locus typicus: Sehnde-Ilten, Lower Saxony, Germany.

Stratum typicum: Bryozoan-rich rubbly limestone in the Belemnitella junior belemnite Zone.

Further distribution: Danian, Limhamn, Skåne län, Sweden.

Stratigraphical range: Late Maastrichtian to Danian.

Fig. 18
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Figure 18: a-b Clavisparsa huckei Voigt, 1962a, holotype, SMF 24141, late Maastrichtian, Neu Wulmstorf-Daerstorf, Lower Saxony, Germany. c-d Mecynoecia (?) latedistans Voigt & Viaud, 1983, holotype, SMF 25488, mid-late Santonian to early Campanian, Les Guignardières in Soullans, Pays de la Loire, France. e-f Frondipora praecursor Voigt, 1951, holotype, SMF 26323, late Maastrichtian (Belemnitella junior belemnite Zone), Sehnde-Ilten, Lower Saxony, Germany.
Scale bars: c 2.5 mm; a, d 500 µm; e 250 µm; b, f 100 µm.

Family Idmoneidae Busk, 1859

Genus Idmonea Lamouroux, 1821

Idmonea rugica (Voigt, 1924)

(Fig. 19a–b )

# 1839 Ceriopora parasitica nob. – Hagenow, p. 286.

# 1846 Ceriopora parasitica v. Hag.Hagenow, p. 594, Pl. XXIII.b, fig. 7.

p# 1899 Reptelea (?) parasitica (von Hagenow), 1839Gregory, p. 294.

*# 1924c Reptotubigera rugica n.sp. – Voigt, p. 5, Pl. I, figs. 11–12.

# 1959a Idmonea parasitica (v. Hagenow), 1839Voigt, p. 39, 50, Pl. IV, figs. 1–2.

Syntypes: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 24157 (Voigt, 1959a, Pl. IV, figs. 1–2).

Original label: Voigt collection number 2434.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Remarks: Voigt (1959a) regarded Idmonea rugica as a synonym of Ceriopora parasitica Hagenow, 1839, based on a specimen found in the Hagenow collection. However, the name Idmonea parasitica is preoccupied by a Recent species from South Australia proposed by Busk (1875). Based on Articles 23.3.5 and 60.2 of the International Code of Zoological Nomenclature (ICZN, 1999), the next oldest available name among the synonyms of I. parasitica is the valid name of this species, which would be Idmonea rugica.

Genus Lagonoecia Canu & Bassler, 1920

Lagonoecia daniensis (Voigt & Eiserhardt, 1995)

(Fig. 19c–d )

*# 1995 Tervidmonea daniensis n.sp. – Voigt & Eiserhardt, p. 419, Figs. 1–10, 25–26.

Holotype: SMF 26132 (Voigt & Eiserhardt, 1995, Figs. 1–4).

Original label: Voigt collection number 13618.

Locus typicus: Stevns Klint in the Stevns Kommune, Zealand Region, Denmark.

Stratum typicum: Hardgrounds of early Danian age.

Further distribution: Early Maastrichtian, chalk pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany. Danian, Agathenburg, Lower Saxony, Germany; Oststeinbek-Havighorst, Schleswig-Holstein, Germany; Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands. Lutetian, Rollgraben near Teisendorf, Bavaria, Germany.

Stratigraphical range: Early Maastrichtian to Lutetian.

Remarks: Tervidmonea daniensis was described as the type species of the new genus Tervidmonea that was placed with reservation in the family Terviidae Canu & Bassler, 1920. However, Viskova (2004b) regarded Tervidmonea as a junior synonym of Lagonoecia Canu & Bassler, 1920.

Family Oncousoeciidae Canu, 1918

Genus Filicisparsa Voigt, 1994

Filicisparsa sommerae Voigt, 1994

(Fig. 19e–f )

# 1991a Genus und Spezies indet. – Voigt, p. 118, Pl. I, figs. 1–2, Pl. II, figs. 1–3.

*# 1994a Filicisparsa sommerae n.g. n.sp. – Voigt, p. 9, Pl. 1, figs. 1–3, Pl. 2, figs. 1–4, Pl. 3, figs. 1–4, Pl. 4, figs. 1–5.

Holotype: SMF 26135 (Voigt, 1994a, Pl. 3, figs. 1–2).

Original label: Voigt collection number 12976.

Locus typicus: Schinkel pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.

Stratum typicum: Micraster rogalae echinoid Zone in the early Santonian.

Further distribution: Early Santonian to late Campanian, Beckum, North Rhine-Westphalia, Germany; Lägerdorf, Schleswig-Holstein, Germany. Campanian, Épernon, Centre-Val de Loire, France; Newhaven, East Sussex and several other localities in the London Basin, England, United Kingdom. Late Campanian, Hacecourt pit near Liège, Wallonia, Belgium. Late Campanian to early Maastrichtian, Saturn pit near Kronsmoor, Schleswig-Holstein, Germany. Late Maastrichtian, Stevns Klint in the Stevns Kommune, Zealand Region, Denmark.

Stratigraphical range: Early Santonian to late Maastrichtian.

Remarks: Filicisparsa sommerae is the type species of Filicisparsa Voigt, 1994. The species was described and figured in a preliminary note (Voigt, 1991a) who recognized it as a representative of a new genus but deferred its introduction pending the finding of specimens with gonozooids.

Fig. 19
Click on thumbnail to enlarge the image.

Figure 19: a-b Idmonea rugica (Voigt, 1924), neotype, SMF 24157, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Lagonoecia daniensis (Voigt & Eiserhardt, 1995), holotype, SMF 26132, early Danian, Stevns Klint in the Stevns Kommune, Zealand Region, Denmark. e-f Filicisparsa sommerae Voigt, 1994, holotype, SMF 26135, early Santonian (Micraster rogalae echinoid Zone), Schinkel pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.
Scale bars: a 2.5 mm; c, e 500 µm; b, d 250 µm; f 100 µm.

Genus Proboscina Audouin, 1826

Proboscina denticulata Voigt, 1982

(Fig. 20a–b )

# 1972 Stomatopora bulbosa Levinsen, 1925Brood, p. 224, Pl. XXVIII, figs. 2–3.

*# 1982b "Proboscina" denticulata n.sp. – Voigt, p. 239, Pl. 7, figs. 1–4.

# 1992a Proboscina denticulata VoigtVoigt, Pl. 5, figs. 6–7.

Holotype: SMF 24888 (Voigt, 1982b, Pl. 7, figs. 1–2).

Original label: Voigt collection number 8162.

Locus typicus: Neu Wulmstorf-Daerstorf, Lower Saxony, Germany.

Stratum typicum: Glacial drift deposit containing flint.

Further distribution: Danian, Faxe quarries, Zealand Region, Denmark and many other localities in Denmark; glacial drifts near Neu Wulmstorf, Lower Saxony, Germany; Sophia Jacoba shaft near Hückelhoven, North Rhine-Westphalia, Germany; Island of Fehmarn, Schleswig-Holstein, Germany. Abundant throughout the Danian of Denmark and Sweden according to Brood (1972).

Stratigraphical range: Danian.

Remarks: Two specimens are labelled with the Voigt collection number 8444, but none with the Voigt collection number 8162. One sample with the Voigt collection number 8444 can be recognized as the holotype.

Proboscina truncata (Hagenow, 1839)

(Fig. 20c–d )

*# 1839 Cellepora truncata nob. – Hagenow, p. 279.

# 1959a Proboscina truncata (v. Hagenow), 1839Voigt, p. 32, Pl. I, fig. 4.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26387 (Voigt, 1959a, Pl. I, fig. 4).

Original label: Voigt collection number 384.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Family Multisparsidae Bassler, 1935

Genus Heterohaplooecia Voigt & Viaud, 1983

Heterohaplooecia monticulifera Voigt & Viaud, 1983

(Fig. 20e–f )

*# 1983 Heterohaplooecia monticulifera n.g. n.sp. – Voigt & Viaud, p. 223, Pl. 2, figs. 1–9.

Holotype: SMF 25236 (Voigt & Viaud, 1983, Pl. 2, figs. 1–3).

Original label: Voigt collection number 9817.

Locus typicus: Les Guignardières, Soullans, Vendée, Pays de la Loire, France.

Stratum typicum: Mid-late Santonian to early Campanian.

Further distribution: Coniacian, Saint-Christophe-sur-le-Nais, Indre-et-Loire, Centre-Val de Loire, France. Santonian to Campanian, La Tonelle, Commequiers, Vendée, Pays de la Loire, France. Campanian, railway station, Soullans, Vendée, Pays de la Loire, France.

Stratigraphical range: Coniacian to Campanian.

Remarks: Heterohaplooecia monticulifera is the type species of Heterohaplooecia Voigt & Viaud, 1983.

Fig. 20
Click on thumbnail to enlarge the image.

Figure 20: a-b Proboscina denticulata Voigt, 1982, holotype, SMF 24888, Danian, Neu Wulmstorf-Daerstorf, Lower Saxony, Germany. c-d Proboscina truncata (Hagenow, 1839), neotype, SMF 26387, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. e-f Heterohaplooecia monticulifera Voigt & Viaud, 1983, holotype, SMF 25236, mid-late Santonian to early Campanian, Les Guignardières in Soullans, Pays de la Loire, France.
Scale bars: c, e 1 mm; a, d 500 µm; f 250 µm; b 100 µm.

Genus Reptoclausa Orbigny, 1853

Reptoclausa wolffi Voigt, 1924

(Fig. 21a–b )

*# 1924a Reptoclausa wolffi n.sp. – Voigt, p. 146, Pl. III, fig. 16.

# 1973a Reptoclausa wolffi Voigt, 1924 – Voigt, p. 141, Pl. 9, figs. 7–8.

Lectotype (defined in Voigt, 1973a): Voigt (1924a, Pl. III, figs. 1–2). This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Burgberg west of Gehrden, Lower Saxony, Germany.

Stratum typicum: (Ober-)Emscher Formation.

Neotype: SMF 26433 (Voigt, 1973a, Pl. 9, fig. 8).

Original label: Voigt collection number 6716.

Locus neotypicus: Brick factory Metje, Burgberg west of Gehrden, Lower Saxony, Germany.

Stratum neotypicum: Late middle Santonian.

Further distribution: Early Cenomanian, Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. Santonian, Sudmerberg near Goslar; abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten; Lengede-Broistedt; Söhlde-Hoheneggelsen (all Lower Saxony, Germany). Early Campanian, Karlshamn, Blekinge län, Sweden.

Stratigraphical range: Early Cenomanian to early Campanian.

Family Plagioeciidae Canu, 1918

Genus Diplosolen Canu, 1918

Diplosolen pavonius Voigt, 1929

(Fig. 21c–d )

# 1839 Cellepora flabelliformis nob. – Hagenow, p. 279.

*# 1929 Diplosolen pavonius n.sp. – Voigt, p. 3, figs. 6–7.

# 1959a Diplosolen pavionus Voigt, 1929Voigt, p. 33, Pl. II, fig. 6.

# 1968a Diplosolen pavionus VoigtVoigt, Pl. I, fig. 1.

Syntypes: This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26391 (Voigt, 1959a, Pl. II, fig. 6).

Original label: Voigt collection number 388.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Further distribution: Early Maastrichtian, Lüneburg-Zeltberg, Lower Saxony, Germany.

Stratigraphical range: Early Maastrichtian.

Remarks: Diplosolen pavionus was considered as a possible junior synonym of Cellepora flabelliformis Hagenow, 1839, by Voigt (1959a). However, the latter was declared as a nomen nudum since the species was not figured by Hagenow and the description was inadequate for its recognition. Two samples labelled Cellepora flabelliformis from the Hagenow collections in Hamburg and Hannover represent two different species, only the specimen from the Hamburg collection belonging to D. pavionus. The species was reported from Lüneburg-Zeltberg by Voigt (1995c).

Genus Plagioecia Canu, 1918

'Plagioecia' disciformis (Hagenow, 1839)

(Fig. 21e–f )

*# 1839 Cellepora disciformis nob. – Hagenow, p. 279.

# 1851 Diastopora disciformis, Hag.Hagenow, p. 16, Pl. X, fig. 7.

p# 1887 Diastopora disciformis v. HagenowMarsson, p. 15.

?# 1899 Berenicea disciformis (Hagenow), 1839Gregory, p. 97, Pl. 6, fig. 2.

# 1959a Berenicea disciformis (v. Hagenow), 1839Voigt, p. 32, Pl. II, fig. 1.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26388 (Voigt, 1959a, Pl. II, fig. 1).

Original label: Voigt collection number 385.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Remarks: The generic assignment of Cellepora disciformis is hampered by the lack of a gonozooid.

Fig. 21
Click on thumbnail to enlarge the image.

Figure 21: a-b Reptoclausa wolffi Voigt, 1924, neotype, SMF 26433, Santonian, Burgberg west of Gehrden, Lower Saxony, Germany. c-d Diplosolen pavonius Voigt, 1929, neotype, SMF 26391, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. e-f Plagioecia disciformis (Hagenow, 1839), neotype, SMF 26388, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany.
Scale bars: c-d 1 mm; a 500 µm; d, f 250 µm; b 100 µm.

'Plagioecia' fistularis (Hagenow, 1846)

(Fig. 22a–b )

*# 1846 Cellepora fistularis v. Hag.Hagenow, p. 612, Pl. XXIII.b, fig. 33.

# 1959a Berenicea fistularis (v. Hagenow), 1839Voigt, p. 54, Pl. II, fig. 5.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26390 (Voigt, 1959a, Pl. II, fig. 5).

Original label: Voigt collection number 387.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Further distribution: Early Maastrichtian, Hemmoor, Lower Saxony, Germany.

Stratigraphical range: Early Maastrichtian.

Remarks: The generic assignment of Cellepora fistularis is hampered by the lack of a gonozooid.

Genus Poroplagioecia Weitschat & Voigt, 1983

Poroplagioecia hartungi Weitschat & Voigt, 1983

(Fig. 22c–d )

*# 1983 Poroplagioecia hartungi n.gen. n.sp. – Weitschat & Voigt, p. 43, Figs. 1–2, Pl. 1, figs. 1–6, Pl. 2, figs. 1–5.

Holotype: SMF 25247 (Weitschat & Voigt, 1983, Pl. 1, figs. 1–6, Pl. 2, figs. 1–5).

Original label: Voigt collection number 7652.

Locus typicus: Brick factory and clay pit K. Gott, Sarstedt, Lower Saxony, Germany.

Stratum typicum: Barremian.

Stratigraphical range: Barremian.

Remarks: Poroplagioecia hartungi is the type species of Poroplagioecia Weitschat & Voigt, 1983. The holotype is the only reported specimen of this species.

Genus Reticrisina Gregory, 1899

Reticrisina crispa Voigt & Viaud, 1983

(Fig. 22e–f )

*# 1983 Reticrisina crispa n.sp. – Voigt & Viaud, p. 222, Pl. 1, figs. 1–6.

Holotype: SMF 25237 (Voigt & Viaud, 1983, Pl. 1, figs. 1–2).

Original label: Voigt collection number 9849.

Locus typicus: Les Guignardières, Soullans, Vendée, Pays de la Loire, France.

Stratum typicum: Mid-late Santonian to early Campanian.

Stratigraphical range: Mid-late Santonian to early Campanian.

Remarks: In the description of the species, 9836 is indicated as the Voigt collection number of the holotype and reference is made to Pl. 2, fig. 1, which depicts the holotype of Heterohaplooecia monticulifera Voigt & Viaud, 1983, instead. According to the figure captions, Voigt collection number 9912 (= SMF 25244), depicted in Pl. 1, fig. 1, should be the holotype. However, Voigt collection number 9849 is depicted in this image, which is also depicted in Pl. 1, fig. 2. Voigt collection number 9912, which is a specimen of Retecrisina crispa, was not figured by Voigt and Viaud (1983), while a specimen labelled with the Voigt collection number 9836 could not be found in the Voigt collection. We therefore consider the specimen with the Voigt collection number 9849 as the holotype of Retecrisina crispa.

Fig. 22
Click on thumbnail to enlarge the image.

Figure 22: a-b 'Plagioecia' fistularis (Hagenow, 1846), neotype, SMF 26390, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. c-d Poroplagioecia hartungi Weitschat & Voigt, 1983, holotype, SMF 25247, Barremian, brick factory and clay pit K. Gott in Sarstedt, Lower Saxony, Germany. e-f Reticrisina crispa Voigt & Viaud, 1983, holotype, SMF 25237, mid-late Santonian to early Campanian, Les Guignardières in Soullans, Pays de la Loire, France.
Scale bars: a, c, e 1 mm; b, d, f 500 µm.

Family Semiceidae Buge, 1952

Genus Filicea Orbigny, 1854

Filicea cincta Voigt, 1924

(Fig. 23a–b )

*# 1924a Filicea cincta n.sp. – Voigt, p. 170, Pl. IV, figs. 3, 6.

# 1973a Filicea cincta Voigt, 1924Voigt, p. 141, Pl. 13, figs. 1–4.

Syntypes: This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Sudmerberg near Goslar and Burgberg west of Gehrden (all Lower Saxony, Germany).

Stratum typicum: Middle Santonian.

Neotype: SMF 24449 (Voigt, 1973a, Pl. 13, fig. 2).

Original label: Voigt collection number 6690.

Locus neotypicus: Burgberg near Gehrden, Lower Saxony, Germany.

Stratum neotypicum: Late Santonian.

Further distribution: Santonian: Ilsede-Groß Bülten; Gehrden; Bad Harzburg; Sudmerberg near Goslar; Abandoned iron-ore mine near Vechelde-Vallstedt (all Lower Saxony, Germany).

Stratigraphical range: Santonian.

Remarks: W. Pockrandt collected the neotype and gave it to Voigt.

Filicea velata (Hagenow, 1839)

(Fig. 23c–d )

# 1839 Ceriopora rhombifera nob. – Hagenow, p. 284.

*# 1839 Ceriopora velata nob. – Hagenow, p. 285, Pl. V, figs. 6a–b.

# 1846 Vaginopora velata v. Hag.Hagenow, p. 602, Pl. XXIII.b, fig. 19.

# 1881 Ceriopora velata HagenowQuenstedt, p. 280, Pl. 153, fig. 109.

# 1887 Filicea velata v. Hagenow sp. – Marsson, p. 46, Pl. IV, fig. 7.

p# 1889 Filicea velata, HagenowPergens, p. 389.

# 1899 Meliceritites (?) rhombifera (von Hagenow), 1839Gregory, p. 348.

non# 1936 Filicea velata von Hagenow, 1839Allègre, p. 96.

non# 1958 Filicea velata von Hagenow, 1839Ducasse, p. 134, Pl. XX, fig. 4, Pl. XXVI, fig. 4.

# 1959a Filicea velata (v. Hagenow), 1839Voigt, p. 38, Pl. III, figs. 4–6.

# 1998 Filicea velata von HagenowBoardman, Figs. 118–120.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Neotype: SMF 26395 (Voigt, 1959a, Pl. III, fig. 4).

Original label: Voigt collection number 392.

Locus neotypicus: Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum neotypicum: White chalk of early Maastrichtian age.

Stratigraphical range: Early Maastrichtian.

Remarks: Voigt (1959a) regarded Ceriopora rhombifera Hagenow, 1839, as a junior synonym of F. velata because intermediate stages between the two putative species exist.

Genus Septocea Voigt, 1995

Septocea septifera Voigt, 1995

(Fig. 23e–f )

*# 1995d Septocea septifera n.g. n.sp. – Voigt, p. 174, Figs. 1–10.

Holotype: SMF 26106 (Voigt, 1995d, Figs. 1, 8).

Original label: Voigt collection number 13476.

Locus typicus: Stubbenkammer, Island of Rügen, Mecklenburg-Vorpommern, Germany.

Stratum typicum: White chalk of early Maastrichtian age.

Further distribution: Late Maastrichtian, Tuffeau de Maastricht (Meerssen Limestone), abandoned van der Zwaan Quarry on the hill Sint-Pietersberg; Curfs Quarry near Berg, Valkenburg aan de Geul municipality (both Limburg, Netherlands).

Stratigraphical range: Maastrichtian.

Remarks: Septocea septifera is the type species of Septocea Voigt, 1995.

Fig. 23
Click on thumbnail to enlarge the image.

Figure 23: a-b Filicea cincta Voigt, 1924, neotype, SMF 24449, late Santonian, Burgberg near Gehrden, Lower Saxony, Germany. c-d Filicea velata (Hagenow, 1839), neotype, SMF 26395, early Maastrichtian, Island of Rügen, Mecklenburg-Vorpommern, Germany. e-f Septocea septifera Voigt, 1995, holotype, SMF 26106, early Maastrichtian, Stubbenkammer on the Island of Rügen, Mecklenburg-Vorpommern, Germany.
Scale bars: a 2.5 mm; c 1 mm; e 500 µm; b 250 µm; d, f 100 µm.

Family Spiroporidae Voigt, 1968

Genus Spiropora Lamouroux, 1821

Spiropora (?) ingens Voigt, 1924

(Fig. 24a–b )

*# 1924a Spiropora ingens n.sp. – Voigt, p. 136, Pl. IV, figs. 17–19.

# 1970 Spiropora (?) ingens Voigt, 1924Voigt & Flor, p. 60, Fig. 24.17, Pl. 15, figs. 6–7.

# 1973a Spiropora (?) ingens Voigt, 1924Voigt, p. 118, Pl. 7, figs. 1–2.

Syntypes: This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Sudmerberg near Goslar and Burgberg west of Gehrden (all Lower Saxony, Germany).

Stratum typicum: Middle Santonian.

Neotype: SMF 24360 (Voigt & Flor, 1970, Pl. 15, fig. 6).

Original label: Voigt collection number 5343.

Locus neotypicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum neotypicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Further distribution: Middle Santonian, Sudmerberg near Goslar; Butterberg near Bad Harzburg; Former building pit on the Ilsenburger Straße, Bad Harzburg; Burgberg west of Gehrden; Söhlde-Hoheneggelsen; Bad Harzburg-Harlingerode (all Lower Saxony, Germany).

Stratigraphical range: Middle Santonian.

Remarks: The neotype originally belonged to the Brandes Collection.

Spiropora suecica Voigt & Flor, 1970

(Fig. 24c–d )

*# 1970 Spiropora suecica n.sp. – Voigt & Flor, p. 57, Figs. 22, 24.13–14, Pl. 8, fig. 6, Pl. 12, figs. 1–10, Pl. 13, figs. 1–2.

non# 1972 Spiropora verticellata (Goldfuss), 1827 [sic] – Brood, p. 309, Pl. XLIII, figs. 6, 12, Pl. LXII, fig. 7.

# 1983 Spiropora suecica Voigt & FlorVoigt, Pl. 5, figs. 16–18.

Holotype: SMF 24351 (Voigt & Flor, 1970, Pl. 12, fig. 2).

Original label: Voigt collection number 5338.

Locus typicus: Hemmingslycke near Kristianstad, Skåne län, Sweden.

Stratum typicum: Late Campanian.

Further distribution: Early Campanian (Belemnellocamax mammillatus belemnite Zone), Karlshamn and Blekinge, Blekinge län, Sweden; Balsberg north of Kristianstad, Skåne län, Sweden; "Ifö" (probably the island Ivön, formerly spelled Ifön near Bromölla, Skåne län, Sweden). Late Campanian, Staversvad near Kristianstad-Arkelstorp; Båstad (both Skåne län, Sweden).

Stratigraphical range: Campanian.

Remarks: Although this species was regarded as a junior synonym of Spiropora verticellata (Goldfuß, 1826) by Brood (1972), Voigt (1973a) confirmed that the two species differ, especially in the morphology of the gonozooids.

Family Theonoidae Busk, 1859

Genus Fascigera Voigt, 1967

Fascigera fasciculus Voigt, 1967

(Fig. 24e–f )

*# 1967 Fascigera fasciculus n.sp. – Voigt, p. 82, Pl. 31, figs. 1–5, Pl. 32, fig. 6.

Holotype: SMF 24214 (Voigt, 1967, Pl. 31, fig. 4, Pl. 32, fig. 6).

Original label: Voigt collection number 4633.

Locus typicus: Hemmoor, Lower Saxony, Germany.

Stratum typicum: White chalk of the late Maastrichtian.

Further distribution: Maastrichtian, unspecified locality along the Emba River (Kazakh: Ембі or Жем; Russian: Эмба) in the Aktobe or Mangystau Province, Kazakhstan.

Stratigraphical range: Maastrichtian.

Remarks: Fascigera fasciculus is the type species of Fascigera Voigt, 1967. In the species description, 4630 is indicated as the Voigt collection number of the holotype, while in the figure captions, both 4623 (Pl. 31, fig. 4) and 4633 (Pl. 32, fig. 6) are indicated as the Voigt collection numbers of the holotype. According to the original label, Voigt collection number 4633 has to be regarded as the holotype of F. fasciculus. Voigt collection number 4630 could not be found and Voigt collection number 4623 is an unimaged sample of F. fasciculus. Fascigeridae was not based on a type genus when Orbigny (1853, p. 665) created this family and all genera included by Orbigny (1853) in the family have since been placed in other families. Many authors have therefore considered the name Fascigeridae as invalid (e.g., Bassler, 1953). Canu (1916, p. 332) used the genus name Fascigera when he assigned Defrancia dimidiata Reuss, 1848 (now included in the genus Tubulipora Lamarck, 1816), to it, but provided no description of the genus and no type species was selected. Fascigera was therefore not a valid generic name at this time. Voigt (1967), who was aware of these problems, established a genus Fascigera to validate the family name Fascigeridae with Orbigny (1853) as its author, assigning Defrancia pluma Reuss, 1848, to it. The legitimacy of this subsequent validation of a previously invalid nominal family taxon was questioned by Walter (1986, p. 82). The International Code on Zoological Nomenclature clearly states that a new family name cannot be based on a non-existent generic name (ICZN, 1999, Article 11.7.1.1.). Therefore, Orbigny (1853) cannot be considered as the author of Fascigeridae. Furthermore, the necessity for a family Fascigeridae is questionable and Fascigera is therefore transferred to Theonoidae Busk, 1859, a family in which F. fasciculus readily fits.

Fig. 24
Click on thumbnail to enlarge the image.

Figure 24: a-b Spiropora (?) ingens Voigt, 1924, neotype, SMF 24360, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany. c-d Spiropora suecica Voigt & Flor, 1970, holotype, SMF 24351, late Campanian, Hemmingslycke near Kristianstad, Skåne län, Sweden. e-f Fascigera fasciculus Voigt, 1967, holotype, SMF 24214, late Maastrichtian, Hemmoor, Lower Saxony, Germany.
Scale bars: a, c 1 mm; b, d-e 500 µm; f 250 µm.

Genus Lopholepis Hagenow, 1851

Lopholepis irregularis Hagenow, 1851

(Fig. 25a–b )

*# 1851 Lopholepis irregularis, Hag.Hagenow, p. 39, Pl. III, figs. 11h–k.

# 1853 Lopholepis irregularis, de Hagenow, 1851Orbigny, p. 687.

# 1857 Lopholepis irregularis, d'Orb.Pictet, Pl. XCI, fig. 6.

p# 1881 Lopholepis radians v. Hag.Hamm, p. 32.

# 1885 Lopholepis rapax, sp. n. – Meunier & Pergens, p. 33, Pl. II, fig. 6.

p# 1909 Lopholepis radians, von Hagenow, 1851Gregory, p. 50.

# 1982b Lopholepis irregularis v. Hagenow, 1851Voigt, p. 63, Pl. 10, figs. 1–3, Pl. 11, figs. 1–6, Pl. 12, figs. 1–5, Pl. 13, figs. 1–5.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Maastricht, Limburg, Netherlands.

Stratum typicum: Tuffeau de Maastricht.

Neotype: SMF 24914 (Voigt, 1982a, Pl. 12, fig. 1).

Original label: Voigt collection number 9314.

Locus neotypicus: Locality near Maastricht, Limburg, Netherlands (see Remarks for further information).

Further distribution: Late Maastrichtian, Curfs Quarry near Berg and Blom Quarry near Terblijt (both in Valkenburg aan de Geul municipality); abandoned van der Zwaan Quarry and ENCI (Eerste Nederlandse Cement Industrie) pit on the hill Sint-Pietersberg (all Limburg, Netherlands); Albert Canal near the Château Neercanne; Riemst-Kanne (both Flanders, Belgium).

Stratum neotypicum: Tuffeau de Maastricht.

Stratigraphical range: Late Maastrichtian.

Remarks: It is uncertain where the neotype was collected because the specimen is unlabelled. Furthermore, the description of the neotype gives the Albert Canal near Neercanne as the provenance, whereas in the figure captions Blom Quarry is indicated and a note accompanying the neotype mentions the Curfs Quarry.

Lopholepis radians Hagenow, 1851

(Fig. 25c–d )

*# 1851 Lopholepis radians, Hag.Hagenow, p. 39, Pl. III, figs. 11a–c.

# 1851 Cavaria ramosa, Hag.Hagenow, p. 53, Pl. VI, fig. 1.

# 1853 Lopholepis radians, de Hagenow, 1851Orbigny, p. 687.

# 1853 Cavaria irregularis, de Hagenow, 1851Orbigny, p. 797.

# 1881 Cavaria ramosa v. Hag.Hamm, p. 25.

p# 1881 Lopholepis radians v. Hag.Hamm, p. 32.

non# 1887 Cavarinella ramosa v. Hagenow sp. – Marsson, p. 19, Pl. I, fig. 6.

# 1899 Cavaria ramosa, von Hagenow, 1851Gregory, p. 399, Fig. 54.

p# 1909 Lopholepis radians, von Hagenow, 1851Gregory, p. 50.

non# 1972 Reptomulticlausa ramosa (Hagenow), 1887Brood, p. 207, Pl. XXI, figs. 5–8, Pl. LXII, fig. 9.

# 1975b Cavarinella ramosa (v. Hagenow) – Voigt, Pl. 7, figs. 1–6.

# 1982b Lopholepis radians v. Hagenow, 1851Voigt, p. 56, Pl. 1, figs. 1–4, Pl. 2, figs. 1–5, Pl. 3, figs. 1–5, Pl. 4, figs. 1–5, Pl. 5, figs. 1–6, Pl. 6, figs. 1–7, Pl. 7, figs. 1–6, Pl. 8, figs. 1–8, Pl. 9, figs. 1–6.

# 1987c Lopholepis radians v. HagenowVoigt, Fig. 6C.

# 1991 Lopholepis radians v. Hagenow, 1851Schäfer, Pl. 54, fig. 3.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Maastricht, Limburg, Netherlands.

Stratum typicum: Tuffeau de Maastricht.

Neotype: SMF 24944 (Voigt, 1982a, Pl. 1, fig. 2).

Original label: Voigt collection number 9250.

Locus neotypicus: Blom Quarry near Terblijt, Valkenburg aan de Geul municipality, Limburg, Netherlands.

Stratum neotypicum: Tuffeau de Maastricht.

Further distribution: Maastrichtian, Saint-Symphorien, Mons municipality, Wallonia, Belgium; Sehnde-Ilten, Lower Saxony, Germany; Tuffeau de Maastricht of the Maastricht area, Netherlands.

Stratigraphical range: Maastrichtian.

Remarks: Lopholepis radians is the type species of Lopholepis Hagenow, 1851.

Family Tubuliporidae Johnston, 1838

Genus Coronidmonea Voigt, 1975

Coronidmonea coronata Voigt, 1975

(Fig. 25e–f )

*# 1975b Coronidmonea coronata n.g. n.sp. – Voigt, p. 88, Pl. 5, figs. 5–14.

# 1994b Coronidmonea coronata VoigtVoigt, Fig. 2G–M.

Holotype: SMF 24606 (Voigt, 1975b, Pl. 5, figs. 5–7).

Original label: Voigt collection number 3991.

Locus typicus: Stevns Klint in the Stevns Kommune, Zealand Region, Denmark.

Further distribution: Danian, Faxe quarries, Zealand Region and many other localities in Denmark; glacial drift deposits in northern Germany.

Stratum typicum: Bryozoan limestone.

Stratigraphical range: Danian.

Remarks: Coronidmonea coronata is the type species of Coronidmonea Voigt, 1975. The holotype is mistakenly mentioned given in the figure captions as Voigt collection number 3891 instead of 3991.

Fig. 25
Click on thumbnail to enlarge the image.

Figure 25: a-b Lopholepis irregularis Hagenow, 1851, neotype, SMF 24914, late Maastrichtian, locality near Maastricht, Limburg, Netherlands. c-d Lopholepis radians Hagenow, 1851, neotype, SMF 24944, Maastrichtian, Blom Quarry near Terblijt in the municipality Valkenburg aan de Geul, Limburg, Netherlands. e-f Coronidmonea coronata Voigt, 1975, holotype, SMF 24606, Danian, Stevns Klint in the Stevns Kommune, Zealand Region, Denmark.
Scale bars: a, c, e 1 mm; b, d, f 500 µm.

Genus Crisisina Orbigny, 1850

Crisisina conjuncta Voigt, 1987

(Fig. 26a–b )

*# 1987b Crisisina conjuncta n.sp. – Voigt, p. 47, Figs. 2A–C, 3A–H.

Holotype: SMF 25493 (Voigt, 1987b, Fig. 2 A–B).

Original label: Voigt collection number 6130.

Locus typicus: Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands.

Stratum typicum: Belemnitella junior belemnite Zone, Tuffeau de Maastricht (Meerssen Limestone).

Further distribution: Late Maastrichtian, Blom Quarry near Terblijt, Valkenburg aan de Geul municipality; Sint-Pietersberg near Maastricht (both Limburg, Netherlands).

Stratigraphical range: Late Maastrichtian.

Genus Decurella Mongereau & Braga, 1967

Decurella? arnaudae Voigt, 1985

(Fig. 26c–d )

*# 1985a Decurella (?) arnaudae nov. sp. – Voigt, p. 625, Pl. 1, figs. 15–20.

Holotype: SMF 25362 (Voigt, 1985a, Pl. 1, figs. 17–18).

Original label: Voigt collection number 8982.

Locus typicus: Bois de Gareau near Écommoy, Pays de la Loire, France.

Stratum typicum: Late Turonian.

Paratypes: SMF 25359, 25360.

Further distribution: Coniacian to Santonian: Gorges du Nant near Cognin-les-Gorges, Auvergne-Rhône-Alpes, France.

Stratigraphical range: Turonian to Coniacian/Santonian.

Remarks: The assignment of this species to Decurella Mongereau & Braga, 1967, is very uncertain, especially because of the huge stratigraphical gap between the only two species assigned to the genus, with the type species, D. toarensis Mongereau & Braga, 1967, being known only from the Priabonian (Eocene) of Italy.

Genus Pergensella Gregory, 1899

Pergensella geniculata (Hagenow, 1851)

(Fig. 26e–f )

p# 1826 Retepora clathrata nobis – Goldfuß, p. 27, Pl. 9, fig. 12e-f (non Pl. 9, fig. 12a–d).

*# 1851 Idmonea geniculata, Hag.Hagenow, p. 33, Pl. III, fig. 5.

# 1899 Pergensella geniculata (von Hagenow), 1851Gregory, p. 209.

# 1909 Pergensella geniculata (Hag.) – Gregory: Pl. IX, fig. 5.

# 1951 Pergensella geniculata (v. Hagenow, 1851) – Voigt, p. 40, Fig. 6, Pl. 3, figs. 1–2.

# 1953 Pergensella geniculata Hag., 1851Bassler, p. G48, Fig. 16.11.

# 1974 Pergensella geniculata (v. Hagenow, 1851) – Weitschat, p. 63, Fig. 2, Pl. 6, figs. 1–6, Pl. 7, figs. 1–5, Pl. 8, figs. 1–5.

Holotype: The originals of the Hagenow collection in the Stettiner Museum were lost during World War II.

Locus typicus: Maastricht, Limburg, Netherlands.

Stratum typicum: Tuffeau de Maastricht.

Neotype: SMF 26481 (Weitschat, 1974, Pl. 6, figs. 4–6).

Original label: Voigt collection number 6618.

Locus neotypicus: ENCI (Eerste Nederlandse Cement Industrie) pit on the hill Sint-Pietersberg, Limburg, Netherlands.

Stratum neotypicum: Tuffeau de Maastricht (Md).

Further distribution: Early late Maastrichtian, Hemmoor, Lower Saxony, Germany. Late Maastrichtian, Albert Canal near the Château Neercanne, Riemst-Kanne, Flanders, Belgium; Sehnde-Ilten; Glacial drift deposits with flint near Neu Wulmstorf-Daerstorf (both Lower Saxony, Germany); glacial drift deposits on the Segrahner Berg near Gudow; glacial drift deposits on the island of Fehmarn (both Schleswig-Holstein, Germany); glacial drift deposits with white chalk near Hamburg-Hummelsbüttel, Hamburg, Germany; abandoned Schunk Quarry near Voerendaal-Kunrade; Curfs Quarry near Berg, Valkenburg aan de Geul municipality (both Limburg, Netherlands).

Stratigraphical range: Late Maastrichtian.

Remarks: Pergensella geniculata is the type species of Pergensella Gregory, 1899.

Fig. 26
Click on thumbnail to enlarge the image.

Figure 26: a-b Crisisina conjuncta Voigt, 1987, holotype, SMF 25493, late Maastrichtian (Belemnitella junior belemnite Zone), Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands. c-d Decurella (?) arnaudae Voigt, 1985, holotype, SMF 25362, late Turonian, Bois de Gareau near Écommoy, Pays de la Loire, France. e-f Pergensella geniculata (Hagenow, 1851), neotype, SMF 26481, late Maastrichtian, ENCI (Eerste Nederlandse Cement Industrie) pit on the hill Sint-Pietersberg, Limburg, Netherlands.
Scale bars: a 2.5 mm; e 1 mm; c, f 500 µm; b, d 250 µm.

Cyclostomata incertae sedis

Genus Aggregopora Voigt, 1982

Aggregopora catena Voigt, 1982

(Fig. 27a–b )

*# 1982b Aggregopora catena n.g.n.sp. – Voigt, p. 235, Pl. 5, figs. 1–2.

Holotype: SMF 24895 (Voigt, 1982b, Pl. 5, fig. 1).

Original label: Voigt collection number 6881.

Locus typicus: Båstad, Skåne län, Sweden.

Stratum typicum: Rubbly limestone from the Belemnitella mucronata belemnite Zone.

Stratigraphical range: Late Campanian.

Aggregopora catenoides Voigt, 1982

(Fig. 27c–d )

*# 1982b Aggregopora catenoides n.g.n.sp. – Voigt, p. 236, Pl. 6, fig. 3.

Holotype: SMF 24894 (Voigt, 1982b, Pl. 6, fig. 3).

Original label: Voigt collection number 6879.

Locus typicus: Båstad, Skåne län, Sweden.

Stratum typicum: Rubbly limestone from the Belemnitella mucronata belemnite Zone.

Stratigraphical range: Late Campanian.

Remarks: The holotype is the only reported specimen of this species.

Aggregopora prescheri Voigt, 1982

*# 1982b Aggregopora prescheri n.g.n.sp. – Voigt, p. 237, Pl. 6, figs. 4–8.

Holotype: Not found (Voigt, 1982b, Pl. 6, figs. 7–8).

Locus typicus: Hoher Stein, Dresden-Plauen, Saxony, Germany.

Stratum typicum: Rocky shore facies of the Dölzschen Formation in the Actinocamax plenus ammonite Zone (latest Cenomanian).

Stratigraphical range: Latest Cenomanian.

Original label: Voigt collection number 6940.

Remarks: The holotype was listed by Eiserhardt (1998).

Aggregopora verruca Voigt, 1982

(Fig. 27e–f )

*# 1982b Aggregopora verruca n.g.n.sp. – Voigt, p. 236, Pl. 6, figs. 1–2.

Holotype: SMF 24896 (Voigt, 1982b, Pl. 6, figs. 1–2).

Original label: Voigt collection number 2951.

Locus typicus: Abandoned iron ore pit Mathilde near Lengede-Broistedt, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Stratigraphical range: Middle Santonian.

Remarks: The holotype is the only reported specimen of this species.

Fig. 27
Click on thumbnail to enlarge the image.

Figure 27: a-b Aggregopora catena Voigt, 1982, holotype, SMF 24895, late Campanian (Belemnitella mucronata belemnite Zone), Båstad, Skåne län, Sweden. c-d Aggregopora catenoides Voigt, 1982, holotype, SMF 24894, late Campanian (Belemnitella mucronata belemnite Zone), Båstad, Skåne län, Sweden. e-f Aggregopora verruca Voigt, 1982, holotype, SMF 24896, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned iron ore pit Mathilde near Lengede-Broistedt, Lower Saxony, Germany.
Scale bars: a 2.5 mm; c, e 1 mm; d 500 µm; f 250 µm; b 100 µm.

Genus Apatotervia Voigt, 1975

Apatotervia minuta Voigt, 1975

(Fig. 28a–b )

*# 1975a Apatotervia minuta n.g. n.sp. – Voigt, p. 241, Fig. 1, Pl. 2, figs. 1–13.

# 1991a Apathotervia [sic] minuta VoigtVoigt, Pl. II, figs. 4–5.

# 1994a Apathotervia [sic] minuta VoigtVoigt, Pl. 2, fig. 5, Pl. 3, figs. 5–7.

Holotype: SMF 24580 (Voigt, 1975a, Pl. 2, fig. 4).

Original label: Voigt collection number 7432.

Stratum typicum: White chalk of early late Campanian age.

Locus typicus: Chalk pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany.

Further distribution: Early to Middle Santonian, Shaft Staffhorst 1 of the Barbara-Erzbergbau GmbH near Staffhorst, Lower Saxony, Germany. Early Campanian, Sehnde-Höver, Lower Saxony, Germany. Late Campanian, Hannover-Misburg, Lower Saxony, Germany. Latest Campanian, Lüneburg-Zeltberg, Lower Saxony, Germany. Early Maastrichtian, Tornesch, Schleswig-Holstein, Germany; Trimingham, Norfolk, England, United Kingdom.

Stratigraphical range: Early Santonian to early Maastrichtian.

Remarks: Apatotervia minuta is the type species of Apatotervia Voigt, 1975. In later papers, Voigt (1991a, 1994a) misspelled the generic name as Apathotervia.

Genus Bueltenopora Hillmer et al., 1997

Bueltenopora hustedti (Voigt, 1924)

(Fig. 28c–d )

*# 1924a Fasciculipora Hustedti n.sp. – Voigt, p. 122, Pl. III, figs. 1–5.

# 1957b Fasciculipora hustedti Voigt, 1924Voigt, p. 4, Pl. 2, fig. 1.

# 1972 Fasciculipora hustedti VoigtFlor, Pl. 9, fig. 1.

# 1983 "Fasciculipora" hustedti VoigtVoigt, Pl. 5, figs. 8–11.

# 1995 Bueltenopora hustedti (Voigt) – Scholz & Hillmer, Pl. 28, fig. 2.

# 1997 Bueltenopora hustedti (Voigt, 1924) – Hillmer et al., p. 205, Pl. 1, figs. 1–10, Pl. 2, figs. 1–4.

Lectotype (defined in Hillmer et al., 1997): Voigt (1924a, Pl. III, figs. 1–2). This material belonged to the first Voigt Collection that was destroyed in a fire at the Geologisches Staatsinstitut Hamburg in 1943.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Neotype: SMF 26189 (Hillmer et al., 1997, Pl. 1, figs. 1a–c).

Original label: Voigt collection number 14833.

Locus neotypicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum neotypicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Further distribution: Santonian, Lahstedt-Adenstedt; Lengede-Broistedt; Abandoned iron-ore mine near Vechelde-Vallstedt; abandoned Barbara pit near Lengede-Barbecke; Söhlde-Hoheneggelsen; Sudmerberg near Goslar; Bad Harzburg-Harlingerode; Goslar-Oker (all Lower Saxony, Germany). Campanian, abandoned iron-ore mine near Damme (Dümmer), Lower Saxony, Germany. Late Maastrichtian, Saint-Symphorien, Mons municipality, Wallonia, Belgium.

Stratigraphical range: Santonian to late Maastrichtian.

Remarks: Bueltenopora hustedti is the type species of Bueltenopora Hillmer et al., 1997.

Bueltenopora woerdemanni Hillmer et al., 1997

(Fig. 28e–f )

*# 1997 Bueltenopora woerdemanni n.g. n.sp. – Hillmer et al., p. 207, Pl. 3, figs. 1–4.

Locus typicus: Ignaberga, Hässleholms kommun, Skåne län, Sweden.

Stratum typicum: Early Campanian.

Stratigraphical range: Early Campanian.

Original label: Voigt collection number 14803.

Holotype: SMF 26194 (Hillmer et al., 1997, Pl. 3, figs. 1–4).

Remarks: J. Wördemann collected the holotype and gave it to Voigt.

Fig. 28
Click on thumbnail to enlarge the image.

Figure 28: a-b Apatotervia minuta Voigt, 1975, holotype, SMF 24580, early late Campanian, chalk pit of the cement factory Alsen-Breitenburg near Lägerdorf, Schleswig-Holstein, Germany. c-d Bueltenopora hustedti (Voigt, 1924), neotype, SMF 26189, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany. e-f Bueltenopora woerdemanni Hillmer et al., 1997, holotype, SMF 26194, early Campanian, Ignaberga in the Hässleholms kommun, Skåne län, Sweden.
Scale bars: c, e 5 mm; a, d, f 500 µm; b 250 µm.

Genus Fasciculiporina Voigt, 1973

Fasciculiporina pockrandti Voigt, 1973

(Fig. 29a–b )

*# 1973a Fasciculiporina pockrandti n.g. n.sp. – Voigt, p. 132, Pl. 10, figs. 1–6.

Holotype: SMF 24434 (Voigt, 1973a, Pl. 10, figs. 1–2).

Original label: Voigt collection number 6701.

Locus typicus: Burgberg west of Gehrden, Lower Saxony, Germany.

Stratum typicum: Late Santonian.

Further distribution: Santonian, Söhlde-Hoheneggelsen, Lower Saxony, Germany.

Stratigraphical range: Santonian.

Remark: Fasciculiporina pockrandti is the type species of Fasciculiporina Voigt, 1973. W. Pockrandt collected the holotype and gave it to Voigt.

Genus Hemiseptopora Voigt, 1993

Hemiseptopora dentifera Voigt, 1993

*# 1993a Hemiseptopora dentifera n. gen. n.sp. – Voigt, p. 364, Pl. 1, figs. 9–12.

Holotype: Not found (Voigt, 1993a, Pl. 1, figs. 9–12).

Original label: Voigt collection number 11514.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: Voigt (1993a) reported three fragments of Hemiseptopora dentifera, none of which could be found. The holotype was listed by Eiserhardt (1998).

Genus Lunariopsis Voigt, 1993

Lunariopsis cava Voigt, 1993

(Fig. 29c–d )

*# 1993a Lunariopsis cava n. gen. n.sp. – Voigt, p. 366, Pl. 2, figs. 1–8.

Holotype: SMF 25977 (Voigt, 1993a, Pl. 2, figs. 1–5).

Original label: Voigt collection number 10596.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: Lunariopsis cava is the type species of Lunariopsis Voigt, 1993.

Genus Pseudokololophos Martha et al., 2014

Pseudokololophos radioporides Martha et al., 2014

(Fig. 29e–f )

*# 2014 Pseudokololophos radioporides sp. nov. – Martha et al., p. 174, Figs. 6A–F, 7A–C.

Holotype: SMF 30001 (Martha et al., 2014, Figs. 6A–B, 7C).

Original label: KlOvi4.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Paratypes: SMF 30002–30007.

Remarks: Pseudokololophos radioporides is the type species of Pseudokololophos Martha et al., 2014.

Fig. 29
Click on thumbnail to enlarge the image.

Figure 29: a-b Fasciculiporina pockrandti Voigt, 1973, holotype, SMF 24434, late Santonian, Burgberg west of Gehrden, Lower Saxony, Germany. c-d Lunariopsis cava Voigt, 1993, holotype, SMF 25977, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. e-f Pseudokololophos radioporides Martha et al., 2014, holotype, SMF 30001, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: c, e 1 mm; a 500 µm; b, d 250 µm; f 100 µm.

Genus Ramofilisparsa Voigt, 1989

Ramofilisparsa walteri Voigt, 1989

(Fig. 30a–b )

*# 1989a Ramofilisparsa walteri n.g. n.sp. – Voigt, p. 90, Pl. 1, figs. 1–6, Pl. 2, figs. 1–8.

Holotype: SMF 25807 (Voigt, 1989a, Pl. 1, fig. 2, non Pl. 2, fig. 3).

Original label: Voigt collection number 11610.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian).

Stratigraphical range: Earliest Cenomanian.

Remarks: Ramofilisparsa walteri is the type species of Ramofilisparsa Voigt, 1989. In the figure captions, it is indicated that that a lateral view of the holotype is depicted in Pl. 2, fig. 3. However, this figure is not the holotype and it remains unclear which specimen it is.

Genus Retecavella Voigt, 1987

Retecavella broodi Voigt, 1987

(Fig. 30c–d )

*# 1987a Retecavella broodi n.g. n.sp. – Voigt, p. 34, Pl. 6, figs. 1–14, Pl. 8, fig. 11.

Holotype: SMF 25519 (Voigt, 1987a, Pl. 6, figs. 1–2).

Original label: Voigt collection number 2703.

Locus typicus: Curfs Quarry near Berg, Valkenburg aan de Geul municipality, Limburg, Netherlands.

Further distribution: Danian, Mons borehole and Ciply, Mons municipality, Wallonia, Belgium; Albert Canal near Riemst-Vroenhoven, Flanders, Belgium; Neu Wulmstorf-Daerstorf, Lower Saxony, Germany; Beatrix borehole near Neer, Leudal municipality, Limburg, Netherlands; several localities in Denmark and southern Sweden.

Stratum typicum: Pockets in the hardground of the Houthem limestone above the Tuffeau de Maastricht.

Stratigraphical range: Danian.

Remarks: Retecavella broodi is the type species of Retecavella Voigt, 1987.

Genus Spirentalophora Voigt, 1968

Spirentalophora brandesi Voigt & Flor, 1970

(Fig. 30e–f )

*# 1970 Spirentalophora brandesi n.sp. – Voigt & Flor, p. 23, Fig. 7, Pl. 2, figs. 1–5, Pl. 15, fig. 8.

Holotype: SMF 24404 (Voigt & Flor, 1970, Pl. 2, fig. 1).

Original label: Voigt collection number 5190.

Locus typicus: Abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.

Stratum typicum: Glauconitic arenaceous limestone of the Gehrden Formation, Gonioteuthis westfalica belemnite Zone, middle Santonian.

Stratigraphical range: Middle Santonian.

Remarks: The holotype originally belonged to the Brandes collection. The image of the holotype is flipped horizontally in Voigt and Flor (1970).

Fig. 30
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Figure 30: a-b Ramofilisparsa walteri Voigt, 1989, holotype, SMF 25807, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany. c-d Retecavella broodi Voigt, 1987, holotype, SMF 25519, Danian, Curfs Quarry near Berg, Valkenburg aan de Geul, Limburg municipality, Netherlands. e-f Spirentalophora brandesi Voigt & Flor, 1970, holotype, SMF 24404, middle Santonian (Gonioteuthis westfalica belemnite Zone), abandoned quarry of the former iron-ore mine of Bülten-Adenstedt near Ilsede-Groß Bülten, Lower Saxony, Germany.
Scale bars: c, e 1 mm; a 500 µm; d, f 250 µm; b 100 µm.

Spirentalophora cristata Voigt & Flor, 1970

(Fig. 31a–b )

*# 1970 Spirentalophora cristata n.sp. – Voigt & Flor, p. 20, Figs. 4, 5, 24.36–37, Pl. 1, figs. 1–6.

Holotype: SMF 24321 (Voigt & Flor, 1970, Pl. 1, fig. 2).

Original label: Voigt collection number 5538.

Locus typicus: Le Breuil near Thoré-la-Rochette, Centre-Val de Loire, France (probably "Coté de Breuil" along the Loire River or the "Rue du Bruil").

Stratum typicum: Sandy, calcareous marl.

Stratigraphical range: Late Turonian.

Spirentalophora frankei Voigt & Flor, 1970

(Fig. 31c–d )

*# 1970 Spirentalophora frankei n.sp. – Voigt & Flor, p. 21, Figs. 6, 24.22–23, Pl. 1, figs. 7–20.

Holotype: SMF 24309 (Voigt & Flor, 1970, Pl. 1, figs. 9–10).

Original label: Voigt collection number 5054.

Locus typicus: Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.

Stratum typicum: Probably the red limestone from the rocky shore facies of the Essen Greensand Formation in the Neostlingoceras carcitanense ammonite Zone (earliest Cenomanian) (see Remarks for further information).

Stratigraphical range: Probably Cenomanian.

Remarks: A. Franke collected all specimens of this species. The exact horizon in the Rauen Quarry, where the specimens were collected is unknown according to Voigt and Flor (1970). However, the reddish colour of the specimens indicates that they probably came from the red limestone of the Essen Greensand Formation.

Fig. 31
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Figure 31: a-b Spirentalophora cristata Voigt & Flor, 1970, holotype, SMF 24321, late Turonian, Le Breuil near Thoré-la-Rochette, Centre-Val de Loire, France (probably "Coté de Breuil" along the Loire River or the "Rue du Bruil"). c-d Spirentalophora frankei Voigt & Flor, 1970, holotype, SMF 24309, early Cenomanian (Neostlingoceras carcitanense ammonite Zone), Kassenberg, Rauen Quarry in Mülheim an der Ruhr-Broich, North Rhine-Westphalia, Germany.
Scale bars: a 1 mm; c 100 µm; b, d 250 µm.

Spirentalophora inconspicua Voigt & Flor, 1970

(Fig. 32a–b )

*# 1970 Spirentalophora inconspicua n.sp. – Voigt & Flor, p. 24, Figs. 8, 24.38–39, Pl. 3, figs. 1–24.

Holotype: Not found (Voigt & Flor, 1970, Fig. 8A, D, Pl. 3, figs. 16–17)

Original label: Voigt collection number 4731.

Locus typicus: Quarry north of Předboj, Central Bohemian Region, Czech Republic.

Stratum typicum: Transgressive sediments on top of crystalline rocks (Actinocamax plenus ammonite Zone, early Turonian).

Stratigraphical range: Early Turonian.

Remarks: The holotype was listed by Eiserhardt (1998). Here, we image specimen SMF 24367 (Voigt collection number 4694), which was figured by Voigt and Flor (1970, Pl. 3, fig. 4).

Genus Talmontipora Voigt & Ernst, 1985

Talmontipora insolita Voigt & Ernst, 1985

(Fig. 32c–d )

*# 1985a Talmontipora insolita n.g. n.sp. – Voigt & Ernst, p. 105, Pl. 1, figs. 1–5, Pl. 2, figs. 1–2, Pl. 3, figs. 1–4, Pl. 4, figs. 1–4, Pl. 5, figs. 1–5, Pl. 6, figs. 1–5, Pl. 7, figs. 1–4.

Holotype: SMF 25405 (Voigt & Ernst, 1985a, Pl. 1, fig. 3, Pl. 2, fig. 1, Pl. 3, fig. 3).

Original label: Voigt collection number 9188.

Locus typicus: Talmont-sur-Gironde, Charente Maritime, Nouvelle-Aquitanie, France.

Stratum typicum: Campanian, probably Barbezieux Formation.

Stratigraphical range: Late Campanian.

Remarks: Talmontipora insolita is the type species of Talmontipora Voigt & Ernst, 1985.

Fig. 32
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Figure 32: a-b Spirentalophora inconspicua Voigt & Flor, 1970, SMF 24367, early Turonian (Actinocamax plenus ammonite Zone), quarry north of Předboj, Central Bohemian Region, Czech Republic. c-d Talmontipora insolita Voigt & Ernst, 1985, holotype, SMF 25405, late Campanian, Talmont-sur-Gironde, Charente-Maritime, Nouvelle-Aquitaine, France.
Scale bars: c 5 mm; a, d 500 µm; b 250 µm.